Ancient mammal catalogue A to Z
An illustrated guide to extinct mammal branches and their evidence
The catalogue follows extinct mammals and close mammaliaform relatives across deep time, from Permian and Triassic synapsids to Ice Age giants. Across changing habitats, some early whales still walked, others became fully marine, and unrelated herbivores independently evolved tonne-scale bodies and elaborate skull structures.
Every card opens a complete profile that separates the preserved fossil from functional inference and artistic reconstruction. Together the profiles compare different diets, habitats and evolutionary experiments across the Permian, Mesozoic and Cenozoic.
A
Aceratherium
Aceratherium incisivum Kaup, 1832
A Late Miocene European rhinoceros known from incomplete type skulls and more complete Höwenegg skeletons.
Open profileAcritohippus
Acritohippus spp.
A three-toed Miocene horse identified through comparative skull and tooth anatomy.
Open profileAdelobasileus
Adelobasileus cromptoni Lucas & Luo, 1993
A Late Triassic cynodont known from one braincase and discussed near the origin of mammaliaforms.
Open profileAdinotherium
Adinotherium ovinum
A medium-sized Patagonian toxodontid known from skull series and near-complete skeletons, with species names revised repeatedly.
Open profileAdunator
Adunator lehmani Russell, 1964; A. amplus Secord, 2008
A Paleocene mammal known mainly from isolated teeth, with disputed North American assignments and uncertain higher relationships.
Open profileAelurodon
Aelurodon ferox and relatives
A Miocene borophagine canid with robust premolars and several successive species.
Open profileAfrohyrax
Afrohyrax championi; A. namibensis
An early Miocene hyrax known from Rusinga skulls and larger Namibian material.
Open profileAlbertogaudrya
Albertogaudrya unica Ameghino, 1901; A.? carahuasensis remains uncertain
An Eocene South American astrapothere known mainly from teeth; a 2025 molar from Chile extends the documented range southward.
Open profileAletodon
Aletodon gunnelli Gingerich, 1977; A. quadravus; A. conardae
A Paleocene North American mammal known chiefly from teeth and jaw fragments; species limits and family placement vary between revisions.
Open profileAlphadon
Alphadon marshi Simpson, 1927
A small Late Cretaceous metatherian best known from isolated molars in western North America.
Open profileAmbulocetus
Ambulocetus natans
An amphibious early whale with weight-bearing limbs, large feet and a cetacean ear region.
Open profileAmebelodon
Amebelodon fricki; A. floridanus (placement debated)
A Miocene proboscidean with a long jaw and paired flattened lower tusks.
Open profileAmerican mastodon
Mammut americanum
A North American browser known from cusped molars, tusks, rare plant remains and isotope evidence for seasonal movement.
Open profileAmphicyon
Amphicyon major and relatives
A large Miocene amphicyonid with powerful limbs and a versatile carnivore dentition.
Open profileAmphiperatherium
Amphiperatherium
A European metatherian genus known mainly from teeth, with a debated boundary from Peratherium.
Open profileAndrewsarchus
Andrewsarchus mongoliensis
A giant-headed Eocene mammal known from one skull and an uncertain body.
Open profileAnkylodon
Ankylodon annectens, A. progressus
A tiny Paleogene insectivore from western North America, known chiefly from teeth and jaw fragments.
Open profileAntepithecus
Antepithecus brachystephanus Ameghino, 1901
A small middle Eocene Patagonian notoungulate whose museum teeth were consolidated into one recognized species.
Open profileApatemys
Apatemys spp.
An Eocene mammal with elongated fingers documented by a rare articulated skeleton.
Open profileApheliscus
Apheliscus spp.
A small Paleogene mammal whose limb bones prompted a debated link to elephant shrews.
Open profileAphelops
Aphelops megalodus and A. mutilus
A hornless North American Miocene rhinoceros whose enamel records browsing and ecological separation from Teleoceras.
Open profileAphronorus
Aphronorus fraudator
A Paleocene mammal known from teeth, jaw fragments and a notable partial cranium of A. orieli.
Open profileArchaeohyrax
Archaeohyrax patagonicus; A. suniensis
Late Oligocene South American typotheres represented by associated skulls, jaws and a large sample of teeth whose wear changes through growth.
Open profileArchaeopithecus
Archaeopithecus rogeri
A small middle-Eocene Patagonian notoungulate whose species history was clarified by a review of more than 200 specimens.
Open profileArchaeotherium
Archaeotherium mortoni
A North American Oligocene entelodont known from skulls, teeth and evidence for varied feeding mechanics.
Open profileArdipithecus
Ardipithecus
Early Ethiopian hominin combining terrestrial bipedalism with climbing and a grasping big toe.
Open profileArfia
Arfia spp.
An early hyaenodont whose teeth trace a broad range across North America, Europe and Asia.
Open profileArgyrohyrax
Argyrohyrax proavus
A Late Oligocene interathere known from teeth, jaws and a few limb bones; taxonomic revisions clarified its debated synonyms.
Open profileArsinoitherium
Arsinoitherium
A massive Afro-Arabian herbivore with two pairs of bony horn cores and unusual ridged teeth.
Open profileAstrapotherium
Astrapotherium magnum
A large Miocene South American herbivore known for its tusks and a nearly complete A. magnum skeleton.
Open profileAustralopithecus
Australopithecus africanus and relatives
A diverse African hominin genus that walked upright while retaining some climbing adaptations.
Open profileB
Basilosaurus
Basilosaurus
A fully marine late Eocene whale with an elongated spine and tiny hind limbs.
Open profileBathygnathus
Dimetrodon borealis (historically Bathygnathus borealis)
A historical name based on one incomplete upper jaw from Prince Edward Island, now referred to Dimetrodon borealis rather than treated as a separate accepted genus.
Open profileBatodon
Batodon tenuis
A small Late Cretaceous eutherian known from a partial Wyoming jaw and scattered teeth.
Open profileBemalambda
Bemalambda nanhsiungensis; B. pachyoesteus; B. crassa
A Paleocene Chinese pantodont documented by skulls and limb bones.
Open profileBisonalveus
Bisonalveus browni
A late Paleocene Wyoming mammal defined by a lower jaw and cheek teeth.
Open profileBunophorus
Bunophorus spp.
An early Eocene artiodactyl distinguished by multi-cusped teeth and known from rare limb fossils.
Open profileC
Cantius
Cantius spp.
An early adapiform primate known from abundant teeth and a partial grasping skeleton.
Open profileCardiolophus
Cardiolophus radinskyi
An early Eocene perissodactyl known from Wyoming skull and dental fossils, with relationships reassessed in 2026.
Open profileCastorocauda
Castorocauda lutrasimilis Ji et al., 2006
A furry Jurassic docodont with a flattened tail, webbed hind foot and evidence of a semi-aquatic ecology.
Open profileCentetodon
Centetodon spp.
A small North American mammal recognised mainly from distinctive cheek teeth and jaw fragments.
Open profileChasicotatus
Chasicotatus
An Argentine Miocene armadillo known chiefly from diagnostic bony armour plates.
Open profileChriacus
Chriacus spp.
An early placental mammal with a rare climbing-adapted skeleton and contrasting locomotor evidence.
Open profileCimolestes
Cimolestes spp.
A Late Cretaceous North American mammal whose revised genus contains two species known chiefly from teeth and jaw fragments.
Open profileCimolodon
Cimolodon spp.
A Cretaceous multituberculate known mainly from specialized teeth, with new associated bones from Mexico.
Open profileCoryphodon
Coryphodon Owen, 1845
Large early Eocene pantodont known from jaws, teeth and partial skeletons, including Arctic records of seasonal feeding.
Open profileCosoryx
Cosoryx spp.
An extinct pronghorn relative recognised by branched horn cores, with uncertain sheath shape and species boundaries.
Open profileCricetodon
Cricetodon spp.
A Miocene cricetodontine rodent whose tooth series reveal multiple lineages.
Open profileCynodontia
Cynodontia Owen, 1861
A mammal-line clade of therapsid synapsids, distinct from dinosaurs and represented today by mammals.
Open profileCyriacotherium
Cyriacotherium argyreum; C. psamminum
A small Paleocene North American mammal whose unusual teeth prompted competing ideas about its relationships.
Open profileD
Deinotherium
Deinotherium
A giant proboscidean with tusks curving down from the lower jaw, known from Africa and Eurasia.
Open profileDesmatolagus
Desmatolagus spp.
An Asian early lagomorph whose wear-sensitive teeth and disputed family placement complicate its history.
Open profileDesmostylus
Desmostylus hesperus
A Miocene coastal mammal with column-built molars and articulated skeletons from Japan.
Open profileDiacodexis
Diacodexis spp.
An early Eocene artiodactyl with a nearly complete skeleton and a specialized ankle.
Open profileDiceratherium
Diceratherium armatum
A North American rhinoceros with paired nasal horn bases and a taxonomic record refined by repeated revisions.
Open profileDidelphodon
Didelphodon vorax Marsh, 1889
A Late Cretaceous metatherian known from a near-complete Hell Creek skull, robust jaws and evidence for a varied diet.
Open profileDidymictis
Didymictis spp.
A viverravid carnivoramorphan whose juvenile teeth corrected an early fossil record.
Open profileDidymoconus
Didymoconus colgatei; D. berkeyi
An enigmatic Oligocene Asian mammal known from skulls, teeth and a Chinese specimen with robust forelimbs.
Open profileDimetrodon
Dimetrodon
A diverse Early Permian predatory synapsid known for its tall neural spines and differentiated teeth.
Open profileDinictis
Dinictis felina
A cat-like North American nimravid distinguished from true cats by skull and ear anatomy.
Open profileDioplotherium
Dioplotherium manigaulti; D. allisoni
A tusked dugongid sirenian known from incomplete fossils in the Americas; the type of D. manigaulti is a premaxillary fragment and tusk.
Open profileDissacus
Dissacus praenuntius
A mesonychid mammal whose Bighorn Basin teeth reveal feeding changes around the PETM.
Open profileDorcatherium
Dorcatherium spp.
A widespread Miocene tragulid with M-structured molars, no antlers and enlarged canines in males.
Open profileDorudon
Dorudon atrox
A five-metre Eocene whale from Fayum with flippers, reduced hind limbs and a rich fossil record at Wadi Al Hitan.
Open profileDusisiren
Dusisiren spp.
Miocene North Pacific sea cows whose California and Japanese fossils preserve changes in teeth and forelimbs.
Open profileDvinia
Dvinia prima
A small Late Permian cynodont from northern Russia, known from skulls with a secondary palate and complex postcanine teeth.
Open profileE
Ectocion
Ectocion spp.
A phenacodontid whose fossil teeth trace body-size and shape patterns across early Cenozoic warming.
Open profileEctypodus
Ectypodus spp.
A multituberculate known mainly from isolated teeth, with rare skull material and an Arctic Canadian occurrence.
Open profileElasmotherium
Elasmotherium sibiricum Fischer, 1808
A giant Pleistocene Eurasian rhinoceros with a frontal skull boss and high-crowned grazing teeth.
Open profileEleutherocercus
Eleutherocercus setifer; E. solidus; E. antiquus
A South American glyptodont studied from armour, skull and tail fossils.
Open profileEllipsodon
Ellipsodon; historical assignments include E. aquilonius and E. priscus
A Paleocene North American mammal known from jaws and teeth, with species names repeatedly transferred or synonymized.
Open profileEntelodon
Entelodon
A long-legged Eurasian entelodont with an enormous head and evidence for an omnivorous diet.
Open profileEntelodonts
Entelodontidae
An extinct family of large artiodactyls known from distinctive skulls, powerful jaws and evidence for varied diets.
Open profileEohyrax
Eohyrax rusticus; E. isotemnoides and relatives
An Eocene South American typothere known mainly from teeth; its species roster and family placement have been revised.
Open profileEomorphippus
Eomorphippus obscurus; E. bondi; E. neilopdykei
A South American notohippid traced through teeth, jaws and revised fossil layers.
Open profileEosclerocalyptus
Eosclerocalyptus lineatus, E. proximus and E. tapinocephalus
Argentine glyptodonts of the Late Miocene and Early Pliocene, known from carapace plates, tail tubes and limb bones.
Open profileEosiren
Eosiren libyca; E. stromeri; other assignments vary by revision
Egyptian Eocene–Oligocene sea cows known from skulls, teeth and postcranial remains; E. libyca preserves a strongly reduced pelvis.
Open profileEotheroides
Eotheroides aegyptiacum
An Eocene sea cow genus with an endocast for its type species and more complete skeletons for later congeners.
Open profileEpicyon
Epicyon saevus, E. haydeni and relatives
A large borophagine canid with massive jaws and premolars capable of processing bone.
Open profileEsthonyx
Esthonyx spp.
An early Eocene tillodont with rooted enlarged incisors and a long, species-rich record in Wyoming.
Open profileEucricetodon
Eucricetodon spp.
An Oligocene cricetid known from diverse tooth samples across Europe and Asia.
Open profileEudinoceras
Eudinoceras mongoliensis and five other recognised species
An Asian Eocene coryphodontid pantodont whose taxonomy and stratigraphic range have been revised through dental comparisons.
Open profileEumyarion
Eumyarion spp.
A Miocene cricetid whose overlapping molar characters complicate species identification.
Open profileEurhinodelphis
Eurhinodelphis spp.
A Miocene toothed whale with an exceptionally elongated, toothless upper rostrum.
Open profileG
Galerix
Galerix spp.
A Miocene galericine known mainly from teeth that document species turnover and regional biochronology.
Open profileGelastops
Gelastops parcus and G. joni
A very small Paleocene mammal known from jaws and teeth in Montana and Wyoming, with a fragmentary referred tooth from Colorado.
Open profileGomphotherium
Gomphotherium angustidens and relatives
A widespread Miocene proboscidean with an elongated lower jaw, upper and lower tusks, and diets that varied among populations.
Open profileGraecopithecus
Graecopithecus freybergi
Late Miocene eastern Mediterranean ape known from a damaged jaw and one premolar, with disputed hominin affinities.
Open profileGranastrapotherium
Granastrapotherium snorki
A giant middle Miocene astrapothere known chiefly from a Colombian mandible and tusks, with an uncertain partial-skull referral from Peru.
Open profileH
Haplaletes
Haplaletes disceptatrix, H. pelicatus; H. serior is disputed
A Paleocene North American mammal known from jaws and isolated teeth, with changing species limits and a debated referral of H. serior.
Open profileHaplomylus
Haplomylus spp.
A small Paleogene mammal whose limb fossils informed a debated hypothesis about elephant-shrew origins.
Open profileHegetotherium
Hegetotherium mirabile; H. cerdasensis; H. novum
A Miocene South American notoungulate with distinctive incisors and a revised, still debated species history.
Open profileHemihegetotherium
Hemihegetotherium achataleptum, H. torresi, H. tantillum and H. trilobus
A Miocene South American hegetotheriid known mainly from teeth and skulls, with rarer limb fossils and a revised species history.
Open profileHenricosbornia
Henricosbornia spp.
An early South American notoungulate known chiefly from Paleogene teeth and jaw fragments.
Open profileHeosminthus
Heosminthus primiveris
A small fossil dipodoid traced through cheek teeth, with species limits and family placement under review.
Open profileHeptodon
Heptodon spp.
An early Eocene tapiroid whose feet and skull preserve an anatomy unlike the modern tapir form.
Open profileHerpetotherium
Herpetotherium cf. fugax
A metatherian with rare Oligocene evidence from the ear region, endocast and hind limb.
Open profileHesperocyon
Hesperocyon gregarius Cope, 1873
An early North American canid combining efficient ground travel with retained climbing features.
Open profileHeterohyus
Heterohyus armatus, H. nanus and related species
An Eocene apatemyid known from European jaws and nearly complete Messel skeletons with elongated fingers.
Open profileHomalodotherium
Homalodotherium cunninghami
A clawed Patagonian notoungulate studied from skulls and limb bones.
Open profileHomo antecessor
Homo antecessor
An early European human from Gran Dolina, known from fragmentary fossils, cut marks and ancient enamel proteins.
Open profileHomo erectus
Homo erectus
A long-lived early human lineage known from Java, Africa and much of Eurasia, with a fully terrestrial body and a fossil record that resists a single tidy origin story.
Open profileHomo ergaster
Homo ergaster
Early African Homo is known from skulls, teeth and a remarkably complete juvenile skeleton, but its species boundary with Homo erectus remains debated.
Open profileHomo floresiensis
Homo floresiensis
A small-bodied island human from Flores, with a long fossil record and unresolved origins.
Open profileHomo georgicus
Homo georgicus
Dmanisi preserves several small-brained early humans in one landscape, offering a rare view of variation during the first dispersals beyond Africa.
Open profileHomo habilis
Homo habilis
Early African human known from a disputed mixture of skull, jaw, hand and limb fossils associated with Oldowan technology.
Open profileHomo heidelbergensis
Homo heidelbergensis
A name founded on one powerful jaw became a label for varied Middle Pleistocene humans, but its geographic and taxonomic limits remain unsettled.
Open profileHomo rudolfensis
Homo rudolfensis
An early human known chiefly from a broad-faced cranium and separate jaws from East Africa.
Open profileHomo sapiens
Homo sapiens
Our species emerged through a long African history: early fossils combine features in different ways, and later populations dispersed and mixed with other humans.
Open profileHomogalax
Homogalax protapirinus
An early Eocene perissodactyl whose partial skeleton preserves primitive limb proportions and unequal digit counts.
Open profileHoplocetus
Hoplocetus ritzi
Miocene physeteroid with an associated German series of teeth and vertebrae; isolated tooth assignments remain debated.
Open profileHuilatherium
Huilatherium pluriplicatum
A Colombian Miocene notoungulate best documented by deciduous teeth; newer Castilletes fossils remain identified only to genus.
Open profileHyaenodon
Hyaenodon leptorhynchus and relatives
A diverse Eocene to Miocene hyaenodont with rear-positioned shearing molars and species-dependent body size.
Open profileHyopsodus
Hyopsodus spp.
A widespread Eocene mammal known from changing tooth populations and one skull that raised a debated hearing hypothesis.
Open profileHypnomys
Hypnomys morpheus
Balearic island dormouse known from cave teeth and articulated skeletons; limb use remains debated.
Open profileHypohippus
Hypohippus spp.
A Miocene three-toed browsing horse known from early tooth finds and later skeletons.
Open profileHyrachyus
Hyrachyus modestus
A middle Eocene perissodactyl known from an unusually informative skeleton with four-toed forefeet and three-toed hind feet.
Open profileHyracodon
Hyracodon spp.
A hornless North American rhinocerotoid whose limb proportions complicate the “running rhino” label.
Open profileHyracotherium
Hyracotherium leporinum
An English early Eocene perissodactyl whose type skull anchors a genus once applied to many different mammals.
Open profileI
Inostrancevia
Inostrancevia alexandri and relatives
A large Late Permian gorgonopsian with blade-like upper canines, known from Russia and later African records.
Open profileInteratherium
Interatherium rodens; I. extensus
A Miocene Patagonian notoungulate whose two accepted species were clarified in a 2023 systematic revision.
Open profileIschyromys
Ischyromys spp.
An early rodent with partial skeletons, a scanned Oligocene skull and a record of species replacement.
Open profileIsotemnus
Isotemnus primitivus and other referred material
A Patagonian Eocene notoungulate known from teeth, jaws and isolated limb bones, with a cautious astragalus-based mass estimate.
Open profileK
Kabwe 1
African Middle Pleistocene Homo
A robust Zambian cranium dated to about 299,000 years ago, with a debated species assignment.
Open profileKraglievichia
Kraglievichia paranensis; K. carinatum
A Miocene pampathere identified mainly from the sculpturing of its armor plates.
Open profileL
Labidolemur
Labidolemur spp.
A North American apatemyid known from type jaws, distinctive teeth and rare cranial and skeletal material.
Open profileLeptauchenia
Leptauchenia decora and relatives
An Oligocene oreodont with high-set eyes, inflated ear chambers and a debated open-country ecology.
Open profileLeptocyon
Leptocyon spp.
Small North American canids that document the long stem history of living Caninae.
Open profileLeptomeryx
Leptomeryx spp.
A small early ruminant known from White River teeth, skulls and limb bones.
Open profileLitaletes
Litaletes disjunctus; related species assignments debated
A small Paleocene mammal from Montana whose jaws and molars define its type species and test later referrals.
Open profileLitocherus
Litocherus zygeus, L. notissimus and L. lacunatus
A Paleocene North American mammal known chiefly from teeth, with an unusual history of competing classifications and some associated postcranial material.
Open profileLitomylus
Litomylus dissentaneus and related Paleocene species
A North American Paleocene mammal known chiefly from teeth, with changing species synonymies and family placement.
Open profileLivyatan
Livyatan melvillei
A giant Miocene sperm whale known from a powerful skull and functional teeth in both jaws.
Open profileLystrosaurus
Lystrosaurus
A widespread dicynodont with a keratinous beak and upper tusks, recorded across the Permian–Triassic boundary.
Open profileM
Maxschlosseria
Maxschlosseria praeterita
A poorly known Patagonian fossil mammal named from upper molars; its species composition and relationships have been repeatedly reviewed.
Open profileMegacerops
Megacerops coloradensis
A massive late Eocene brontothere with paired skull bosses and a long taxonomic history that includes the name Brontotherium.
Open profileMegacricetodon
Megacricetodon spp.
Miocene cricetid rodents whose teeth track regional lineages and dispersal across Europe.
Open profileMegistotherium
Megistotherium osteothlastes
A giant Libyan hyaenodont known chiefly from a 65-centimetre skull and cutting teeth.
Open profileMeniscoessus
Meniscoessus spp.
A Late Cretaceous multituberculate known from incisors, serrated premolars and molars.
Open profileMerychippus
Merychippus sensu lato
Miocene horses with high-crowned teeth and a three-toed foot, within a branching equid radiation.
Open profileMerychyus
Merychyus spp.
A North American oreodont known from changing species, distinctive cheek teeth and rare partial skeletons.
Open profileMerycoidodon
Merycoidodon
A North American oreodont with selenodont plant-processing teeth and a taxonomic history reshaped by abundant fossils.
Open profileMesodma
Mesodma spp.
A small multituberculate known from distinctive premolars in Late Cretaceous and Paleocene deposits.
Open profileMesonyx
Mesonyx obtusidens
A middle Eocene mesonychid known from skulls, teeth and the composite Twin Buttes skeleton.
Open profileMetaxytherium
Metaxytherium spp.
A widespread fossil dugongid with species-specific tusks and skeletons from coastal seas.
Open profileMeteutatus
Meteutatus lagenaformis and related species
A Patagonian Paleogene armadillo known chiefly from armour plates, with uncertain boundaries among several historical species names.
Open profileMiacis
Miacis parvivorus Cope, 1872
The type species of a once broad genus, now placed among early carnivoraforms by anatomical analyses.
Open profileMicrodyromys
Microdyromys koenigswaldi
A widespread fossil dormouse genus known mainly from molars whose species boundaries remain under review.
Open profileMicroparamys
Microparamys spp.
A tiny early Eocene rodent known from detailed teeth and a still-fragmentary body record.
Open profileMicrosyops
Microsyops spp.
A Paleogene mammal whose large fossil sample records cavities and changing tooth form.
Open profileMiniochoerus
Miniochoerus spp.
A North American oreodont known from skulls and skeletons across several late Eocene and Oligocene intervals.
Open profileMioclaenus
Mioclaenus turgidus Cope, 1881
A Paleocene North American mammal documented mainly by jaw fragments and teeth; its type is associated with Torrejonian strata in New Mexico.
Open profileMiohippus
Miohippus annectens and related forms
An Oligocene three-toed horse whose Oregon teeth exposed the pitfalls of naming species from wear.
Open profileMoeritherium
Moeritherium lyonsi, M. trigodon, M. chehbeurameuri
An early North African proboscidean whose tooth wear and enamel isotopes support a partly aquatic ecology.
Open profileMylodon
Mylodon darwinii
A South American ground sloth known from Darwin’s type jaw, Patagonian cave remains, preserved hair and plant-bearing coprolites.
Open profileN
Nakalipithecus
Nakalipithecus nakayamai
Late Miocene Kenyan hominoid known from a partial lower jaw and eleven isolated teeth.
Open profileNeanderthals
Homo neanderthalensis
A distinct human lineage known across Pleistocene Europe and western Asia from fossils, tools and ancient DNA.
Open profileNeoplagiaulax
Neoplagiaulax spp.
A Paleocene multituberculate identified chiefly from incisors, premolars and jaws.
Open profileNesodon
Nesodon imbricatus
A large Early Miocene toxodontid known from skulls and substantial skeletal remains; body mass is estimated at about 550 kg.
Open profileNimravids
Nimravidae
An extinct feliform family with several independent forms of enlarged upper canines, separate from true cats.
Open profileNotharctus
Notharctus spp.
An Eocene adapiform known from skulls, limbs and a foot with a probable grooming claw.
Open profileNotopithecus
Notopithecus adapinus Ameghino, 1897
A small Patagonian Eocene notoungulate known from an associated skeleton, growth-related dental variation and revised species names.
Open profileNotostylops
Notostylops murinus Ameghino, 1897
A Patagonian notoungulate known from teeth and a partial skeleton used to study limb function and body-mass estimates.
Open profileNyctitherium
Nyctitherium spp.
A small Eocene mammal identified from jaws and distinctive teeth.
Open profileO
Oldfieldthomasia
Oldfieldthomasia furcata and historically named congeners
An Eocene South American notoungulate whose species list and family placement have been repeatedly reassessed from dental fossils.
Open profileOrohippus
Orohippus pumilus
A small Eocene equid whose teeth and multi-toed feet illuminate a branching early horse family tree.
Open profileOrrorin
Orrorin tugenensis
A late Miocene Kenyan hominin represented by femora, jaw fragments and evidence of both bipedal loading and climbing.
Open profileOxyaena
Oxyaena spp.
An Eocene oxyaenid predator identified by jaws and a specialized shearing dentition.
Open profileP
Pachyrukhos
Pachyrukhos moyani; P. politus
A small Miocene notoungulate with independently evolved rabbit-like proportions; the modern revision recognizes two species.
Open profilePakicetus
Pakicetus inachus and related pakicetids
A land-capable early whale recognised by its cetacean ear and artiodactyl ankle anatomy.
Open profilePalaeolagus
Palaeolagus haydeni
An early North American lagomorph with two pairs of upper incisors and skulls studied using micro-CT.
Open profilePalaeoloxodon
Palaeoloxodon antiquus and relatives
Pleistocene straight-tusked elephants ranging from mainland giants to independently evolved island dwarfs.
Open profilePalaeoryctes
Palaeoryctes puercensis and related species
A small North American fossil mammal known mainly from teeth, jaws and fragmentary skulls; its species limits and higher classification remain debated.
Open profilePalaeosinopa
Palaeosinopa spp.
A pantolestid known from teeth and rare skeletons, including a juvenile from Fossil Butte.
Open profilePalaeotherium
Palaeotherium spp.
A diverse European Eocene perissodactyl genus known from skulls, teeth and three-toed limbs.
Open profilePaleotomus
Paleotomus senior; P. milleri; P. carbonensis; P. radagasti; P. junior
Paleocene eutherians known mostly from isolated teeth in western North America; species limits and family placement have shifted between studies.
Open profilePantolambda
Pantolambda bathmodon Cope, 1882
A Paleocene pantodont from western North America; dental histology offers rare evidence about early mammalian growth.
Open profileParamys
Paramys Leidy, 1871
Early Eocene rodent with a near-complete skeleton used to study climbing and terrestrial locomotion.
Open profileParanthropus
Paranthropus
Three African hominins shared unusually large cheek teeth, yet their diets and evolutionary relationships were more varied than the old 'nutcracker' label suggests.
Open profileParanyctoides
Paranyctoides sternbergi; P. maleficus; P. quadrans and other referred forms
A rare Late Cretaceous eutherian known from teeth and jaw fragments; species limits and Asian referrals remain disputed.
Open profilePararyctes
Pararyctes pattersoni; P. rutherfordi
Paleocene North American eutherians known mainly from teeth and a partial maxilla; their family placement remains debated.
Open profileParastrapotherium
Parastrapotherium holmbergi
A large Patagonian astrapothere traced through teeth and jaw fragments.
Open profileParutaetus
Parutaetus chicoensis; P. chilensis; P. punaensis; P. oliveirai
An early South American armadillo lineage known chiefly from bony carapace plates.
Open profilePeltephilus
Peltephilus spp.
A South American armored xenarthran with bony projections on its head shield, best documented from Oligocene and Miocene fossil localities.
Open profilePeradectes
Peradectes spp.
A Paleogene metatherian whose taxonomy is built chiefly from tiny molars and jaw fragments.
Open profilePericotoxodon
Pericotoxodon platignathus
A large middle Miocene Colombian toxodontid known from partial skulls, teeth and a femur, with modelled mass estimates and fossil bite traces.
Open profilePeriphragnis
Periphragnis harmeri Roth, 1899
A South American notoungulate represented by separate Eocene and Oligocene records in Patagonia and Chile.
Open profilePeriptychus
Periptychus carinidens
A stout Paleocene mammal with folded teeth, a robust skull and plantigrade limbs.
Open profilePhenacodaptes
Phenacodaptes sabulosus
A late Paleocene Wyoming mammal studied mainly from its jaws and teeth.
Open profilePhenacodus
Phenacodus primaevus Cope, 1873 and relatives
A five-toed early placental whose skeletons document incipient terrestrial running specialisation.
Open profilePhiomia
Phiomia serridens; P. major
An Oligocene early proboscidean with projecting lower incisors.
Open profilePlatybelodon
Platybelodon danovi
A Miocene proboscidean with a broadened lower jaw and paired flattened cutting tusks.
Open profilePlesiadapis
Plesiadapis tricuspidens
A Paleogene plesiadapiform with enlarged incisors, claws and a long tail, near but not demonstrably ancestral to primates.
Open profilePleurostylodon
Pleurostylodon
An Eocene Patagonian notoungulate known from teeth and partial skeletons, including material that corrected a composite reconstruction.
Open profilePliohippus
Pliohippus spp.
A Miocene horse genus tracing the gradual, branching evolution of a single-toed limb.
Open profileProcerberus
Procerberus
A small cimolestid mammal known from jaw fragments and teeth around the Cretaceous–Paleogene boundary.
Open profileProeuphractus
Proeuphractus limpidus
A South American Miocene armadillo whose type species is known from separate armour plates and a rediscovered fixed osteoderm.
Open profileProeutatus
Proeutatus
A Patagonian Miocene armadillo represented by osteoderms, skulls and limb bones; several historical species names are under review.
Open profileProhegetotherium
Prohegetotherium sculptum; additional referrals disputed
A late Oligocene hegetotheriid from Argentina, with a type species based on two fragmentary upper jaws.
Open profilePromioclaenus
Promioclaenus acolytus and related species
A Paleocene placental mammal known chiefly from teeth and jaw fragments; its family placement and species boundaries have been revised.
Open profilePropalaehoplophorus
Propalaehoplophorus australis
An early Miocene Patagonian glyptodont known from armor and cranial remains.
Open profileProstegotherium
Prostegotherium notostylopianum; P. astrifer
An early Eocene armadillo-like cingulate diagnosed mostly from patterned armor plates.
Open profileProtoreodon
Protoreodon spp.
An Eocene oreodont revised through comparisons of skulls, teeth and limbs.
Open profileProtoselene
Protoselene opisthacus, P. ashtoni; some referrals remain uncertain
A Paleocene North American mammal known chiefly from teeth and jaw fragments, with a species list shaped by cautious referrals and revisions.
Open profileProtosiren
Protosiren fraasi; P. sattaensis; P. smithae
Eocene sea cows from Egypt and Pakistan whose partial skeletons preserve hind limbs alongside aquatic adaptations.
Open profilePrototherium
Prototherium spp.
Eocene European sea cows known from several skulls and scattered remains; species boundaries remain under review.
Open profilePrototomus
Prototomus spp.
A small early Eocene hyaenodont whose lost type and later Wyoming fossils complicate its taxonomy.
Open profileProtypotherium
Protypotherium spp.
A South American interatheriid whose Miocene teeth and skulls are reshaping species boundaries.
Open profileProzaedyus
Prozaedyus
A small South American armadillo known from skulls and armor plates, with a changing species taxonomy and younger fragmentary records.
Open profilePseudhipparion
Pseudhipparion spp.
A diverse genus of three-toed North American horses with species-specific dental specializations.
Open profilePseudhyrax
Pseudhyrax eutrachytheroides; P. strangulatus
A South American typothere known chiefly from separate teeth; species boundaries and regional referrals require careful comparison.
Open profilePseudoplohophorus
Pseudoplohophorus orientalis, P. absolutus
A Uruguayan Late Miocene glyptodont known from an associated partial skeleton, armour and a tail tube; some species assignments remain uncertain.
Open profilePtilodus
Ptilodus montanus Cope, 1881
A Paleocene multituberculate known for specialised chewing teeth and rare partial skeletons that inform debates about movement.
Open profilePuercolestes
Puercolestes simpsoni
An early Paleocene cimolestid known from a damaged New Mexico skull and provisionally referred Montana teeth.
Open profilePyrotherium
Pyrotherium romeroi, P. macfaddeni
A large Late Oligocene South American mammal known from incomplete skulls, distinctive teeth and scattered limb bones; its relationships and trunk remain uncertain.
Open profileR
Rhynchippus
Rhynchippus equinus; R. pumilus; R. brasiliensis
A Patagonian notoungulate whose skull reveals tooth and middle-ear anatomy.
Open profileS
Sahelanthropus
Sahelanthropus tchadensis
A roughly seven-million-year-old Chadian ape known for the Toumaï cranium and disputed evidence of bipedalism.
Open profileScirrotherium
Scirrotherium hondaensis; S. antelucanus
A Miocene pampathere from Colombia and Costa Rica whose skull and armor prompted a taxonomic revision.
Open profileSciuravus
Sciuravus spp.
An Eocene sciuravid rodent whose skull and teeth complicate the squirrel comparison.
Open profileSifrhippus
Sifrhippus sandrae
An early equid whose teeth document body-size change during rapid PETM warming.
Open profileSinclairella
Sinclairella dakotensis; S. simplicidens
A rare apatemyid known from a crushed Eocene skull and scattered teeth, including a late Oligocene Florida species.
Open profileSinolagomys
Sinolagomys spp.
An early Central Asian pika relative known from a long, tooth-rich Oligocene–Miocene record.
Open profileStegodon
Stegodon
An African and Asian proboscidean genus with ridged molars, diverse diets and multiple island dwarf lineages.
Open profileStegotherium
Stegotherium tessellatum, S. notohippidensis, S. tauberi
A Miocene Patagonian armadillo with a long snout, reduced teeth and a debated type series.
Open profileStenotatus
Stenotatus patagonicus
A South American fossil armadillo identified chiefly by three-ridged movable osteoderms; its type material and species boundaries are debated.
Open profileT
Teleoceras
Teleoceras major Hatcher, 1894 and relatives
A short-limbed rhinoceros known from many articulated Miocene skeletons at Ashfall in Nebraska.
Open profileTetonius
Tetonius homunculus
A small early Eocene primate from Wyoming whose teeth and new basin records have reshaped its taxonomic story.
Open profileTherocephalians
Therocephalia
A diverse Permian and Triassic therapsid group known for varied skulls, differentiated teeth and a fossil record across Pangaea.
Open profileThomashuxleya
Thomashuxleya externa and T. rostrata
A middle-Eocene Patagonian notoungulate known from an associated skeleton that links the skull with major parts of the limbs and girdles.
Open profileThrinaxodon
Thrinaxodon liorhinus
An Early Triassic cynodont known from articulated fossils and burrow associations in southern Gondwana.
Open profileThryptacodon
Thryptacodon spp.
A fragmentary early placental known for a possible tooth-comb-like arrangement.
Open profileThylacoleo
Thylacoleo carnifex
A Pleistocene Australian marsupial predator with shearing premolars, powerful forelimbs and an unusually rich skeletal record.
Open profileThylacosmilus
Thylacosmilus atrox
A South American sparassodont with ever-growing upper canines and a debated role in hunting and carcass processing.
Open profileThyrohyrax
Thyrohyrax domorictus and related species
An early Oligocene Fayum hyracoid known from a type mandible, later crania and an enigmatic chamber in some jaws.
Open profileTitanoides
Titanoides primaevus
A large Late Paleocene pantodont known mainly from rare teeth and jaw fragments in North America.
Open profileTrachytherus
Trachytherus spegazzinianus, T. alloxus
A South American late Oligocene notoungulate known especially from skulls and teeth; Salla specimens refined its anatomy while leaving genus limits debated.
Open profileTrigonostylops
Trigonostylops wortmani
A Patagonian Eocene astrapothere known from a notable skull, jaw fragments and isolated teeth.
Open profileTrogosus
Trogosus spp.
A large middle Eocene tillodont known from jaws, skulls, limb bones and CT-based cranial studies.
Open profileTypotheriopsis
Typotheriopsis chasicoensis, T. silveyrai
An Argentine mesotheriid with two principal species, a debated premolar diagnosis and newly reported associated skeletal material.
Open profileU
Ugandax
Ugandax coryndonae
Early African bovin documented by Pliocene skulls, horn cores and teeth; its precise relationship to living buffalo remains debated.
Open profileUintatherium
Uintatherium anceps
A heavy North American herbivore with three pairs of skull bosses and long upper canines.
Open profileUstatochoerus
Ustatochoerus spp.
A late oreodont genus with revised species boundaries and increasingly tall cheek teeth.
Open profileUtaetus
Utaetus buccatus
An Eocene South American armadillo known mainly from osteoderms, with a revised type history and a confirmed occurrence in southern Brazil.
Open profileV
Vassallia
Vassallia minuta; V. maxima (disputed assignment)
An Argentine pampathere known from jaws and armor; the famous V. maxima skull has been reassigned in later revisions.
Open profileVetelia
Vetelia puncta, V. perforata and V. gandhii
A Miocene armadillo from Argentina and Chile, known from armour plates and rare skulls; a morphological analysis proposed new relationships.
Open profileVillanyia
Villanyia exilis
Small fossil vole defined from Hungarian molars; revised enamel evidence limits older assignments to the genus.
Open profileViverravus
Viverravus spp.
A small viverravid known mostly from teeth, with one partial skeleton supporting agile climbing in V. acutus.
Open profileX
Xenastrapotherium
Xenastrapotherium christi; X. kraglievichi; X. amazonense; X. chaparralensis; X. aequatorialis
Miocene astrapotheres from tropical South America, known mostly from teeth and jaws; genus limits remain under review.
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