Palaeontological atlas

Ancient mammal catalogue A to Z

An illustrated guide to extinct mammal branches and their evidence

The catalogue follows extinct mammals and close mammaliaform relatives across deep time, from Permian and Triassic synapsids to Ice Age giants. Across changing habitats, some early whales still walked, others became fully marine, and unrelated herbivores independently evolved tonne-scale bodies and elaborate skull structures.

Every card opens a complete profile that separates the preserved fossil from functional inference and artistic reconstruction. Together the profiles compare different diets, habitats and evolutionary experiments across the Permian, Mesozoic and Cenozoic.

In the catalogue249mammal profiles
Collection focusMammal originsfrom early synapsids to later mammals
Time rangePermian to Pleistocenefrom early synapsids to Ice Age mammals
Showing 249 of 249 profiles

A

Catalogue illustration for AceratheriumA

Aceratherium

Aceratherium incisivum Kaup, 1832

A Late Miocene European rhinoceros known from incomplete type skulls and more complete Höwenegg skeletons.

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Catalogue illustration for AcritohippusA

Acritohippus

Acritohippus spp.

A three-toed Miocene horse identified through comparative skull and tooth anatomy.

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Catalogue illustration for AdelobasileusA

Adelobasileus

Adelobasileus cromptoni Lucas & Luo, 1993

A Late Triassic cynodont known from one braincase and discussed near the origin of mammaliaforms.

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Catalogue illustration for AdinotheriumA

Adinotherium

Adinotherium ovinum

A medium-sized Patagonian toxodontid known from skull series and near-complete skeletons, with species names revised repeatedly.

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Catalogue illustration for AdunatorA

Adunator

Adunator lehmani Russell, 1964; A. amplus Secord, 2008

A Paleocene mammal known mainly from isolated teeth, with disputed North American assignments and uncertain higher relationships.

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Catalogue illustration for AelurodonA

Aelurodon

Aelurodon ferox and relatives

A Miocene borophagine canid with robust premolars and several successive species.

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Catalogue illustration for AfrohyraxA

Afrohyrax

Afrohyrax championi; A. namibensis

An early Miocene hyrax known from Rusinga skulls and larger Namibian material.

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Catalogue illustration for AlbertogaudryaA

Albertogaudrya

Albertogaudrya unica Ameghino, 1901; A.? carahuasensis remains uncertain

An Eocene South American astrapothere known mainly from teeth; a 2025 molar from Chile extends the documented range southward.

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Catalogue illustration for AletodonA

Aletodon

Aletodon gunnelli Gingerich, 1977; A. quadravus; A. conardae

A Paleocene North American mammal known chiefly from teeth and jaw fragments; species limits and family placement vary between revisions.

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Catalogue illustration for AlphadonA

Alphadon

Alphadon marshi Simpson, 1927

A small Late Cretaceous metatherian best known from isolated molars in western North America.

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Catalogue illustration for AmbulocetusA

Ambulocetus

Ambulocetus natans

An amphibious early whale with weight-bearing limbs, large feet and a cetacean ear region.

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Catalogue illustration for AmebelodonA

Amebelodon

Amebelodon fricki; A. floridanus (placement debated)

A Miocene proboscidean with a long jaw and paired flattened lower tusks.

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Catalogue illustration for American mastodonA

American mastodon

Mammut americanum

A North American browser known from cusped molars, tusks, rare plant remains and isotope evidence for seasonal movement.

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Catalogue illustration for AmphicyonA

Amphicyon

Amphicyon major and relatives

A large Miocene amphicyonid with powerful limbs and a versatile carnivore dentition.

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Catalogue illustration for AmphiperatheriumA

Amphiperatherium

Amphiperatherium

A European metatherian genus known mainly from teeth, with a debated boundary from Peratherium.

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Catalogue illustration for AndrewsarchusA

Andrewsarchus

Andrewsarchus mongoliensis

A giant-headed Eocene mammal known from one skull and an uncertain body.

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Catalogue illustration for AnkylodonA

Ankylodon

Ankylodon annectens, A. progressus

A tiny Paleogene insectivore from western North America, known chiefly from teeth and jaw fragments.

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Catalogue illustration for AntepithecusA

Antepithecus

Antepithecus brachystephanus Ameghino, 1901

A small middle Eocene Patagonian notoungulate whose museum teeth were consolidated into one recognized species.

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Catalogue illustration for ApatemysA

Apatemys

Apatemys spp.

An Eocene mammal with elongated fingers documented by a rare articulated skeleton.

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Catalogue illustration for ApheliscusA

Apheliscus

Apheliscus spp.

A small Paleogene mammal whose limb bones prompted a debated link to elephant shrews.

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Catalogue illustration for AphelopsA

Aphelops

Aphelops megalodus and A. mutilus

A hornless North American Miocene rhinoceros whose enamel records browsing and ecological separation from Teleoceras.

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Catalogue illustration for AphronorusA

Aphronorus

Aphronorus fraudator

A Paleocene mammal known from teeth, jaw fragments and a notable partial cranium of A. orieli.

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Catalogue illustration for ArchaeohyraxA

Archaeohyrax

Archaeohyrax patagonicus; A. suniensis

Late Oligocene South American typotheres represented by associated skulls, jaws and a large sample of teeth whose wear changes through growth.

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Catalogue illustration for ArchaeopithecusA

Archaeopithecus

Archaeopithecus rogeri

A small middle-Eocene Patagonian notoungulate whose species history was clarified by a review of more than 200 specimens.

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Catalogue illustration for ArchaeotheriumA

Archaeotherium

Archaeotherium mortoni

A North American Oligocene entelodont known from skulls, teeth and evidence for varied feeding mechanics.

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Catalogue illustration for ArdipithecusA

Ardipithecus

Ardipithecus

Early Ethiopian hominin combining terrestrial bipedalism with climbing and a grasping big toe.

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Catalogue illustration for ArfiaA

Arfia

Arfia spp.

An early hyaenodont whose teeth trace a broad range across North America, Europe and Asia.

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Catalogue illustration for ArgyrohyraxA

Argyrohyrax

Argyrohyrax proavus

A Late Oligocene interathere known from teeth, jaws and a few limb bones; taxonomic revisions clarified its debated synonyms.

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Catalogue illustration for ArsinoitheriumA

Arsinoitherium

Arsinoitherium

A massive Afro-Arabian herbivore with two pairs of bony horn cores and unusual ridged teeth.

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Catalogue illustration for AstrapotheriumA

Astrapotherium

Astrapotherium magnum

A large Miocene South American herbivore known for its tusks and a nearly complete A. magnum skeleton.

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Catalogue illustration for AustralopithecusA

Australopithecus

Australopithecus africanus and relatives

A diverse African hominin genus that walked upright while retaining some climbing adaptations.

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B

Catalogue illustration for BasilosaurusB

Basilosaurus

Basilosaurus

A fully marine late Eocene whale with an elongated spine and tiny hind limbs.

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Catalogue illustration for BathygnathusB

Bathygnathus

Dimetrodon borealis (historically Bathygnathus borealis)

A historical name based on one incomplete upper jaw from Prince Edward Island, now referred to Dimetrodon borealis rather than treated as a separate accepted genus.

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Catalogue illustration for BatodonB

Batodon

Batodon tenuis

A small Late Cretaceous eutherian known from a partial Wyoming jaw and scattered teeth.

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Catalogue illustration for BemalambdaB

Bemalambda

Bemalambda nanhsiungensis; B. pachyoesteus; B. crassa

A Paleocene Chinese pantodont documented by skulls and limb bones.

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Catalogue illustration for BisonalveusB

Bisonalveus

Bisonalveus browni

A late Paleocene Wyoming mammal defined by a lower jaw and cheek teeth.

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Catalogue illustration for BunophorusB

Bunophorus

Bunophorus spp.

An early Eocene artiodactyl distinguished by multi-cusped teeth and known from rare limb fossils.

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C

Catalogue illustration for CantiusC

Cantius

Cantius spp.

An early adapiform primate known from abundant teeth and a partial grasping skeleton.

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Catalogue illustration for CardiolophusC

Cardiolophus

Cardiolophus radinskyi

An early Eocene perissodactyl known from Wyoming skull and dental fossils, with relationships reassessed in 2026.

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Catalogue illustration for CastorocaudaC

Castorocauda

Castorocauda lutrasimilis Ji et al., 2006

A furry Jurassic docodont with a flattened tail, webbed hind foot and evidence of a semi-aquatic ecology.

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Catalogue illustration for CentetodonC

Centetodon

Centetodon spp.

A small North American mammal recognised mainly from distinctive cheek teeth and jaw fragments.

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Catalogue illustration for ChasicotatusC

Chasicotatus

Chasicotatus

An Argentine Miocene armadillo known chiefly from diagnostic bony armour plates.

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Catalogue illustration for ChriacusC

Chriacus

Chriacus spp.

An early placental mammal with a rare climbing-adapted skeleton and contrasting locomotor evidence.

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Catalogue illustration for CimolestesC

Cimolestes

Cimolestes spp.

A Late Cretaceous North American mammal whose revised genus contains two species known chiefly from teeth and jaw fragments.

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Catalogue illustration for CimolodonC

Cimolodon

Cimolodon spp.

A Cretaceous multituberculate known mainly from specialized teeth, with new associated bones from Mexico.

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Catalogue illustration for CoryphodonC

Coryphodon

Coryphodon Owen, 1845

Large early Eocene pantodont known from jaws, teeth and partial skeletons, including Arctic records of seasonal feeding.

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Catalogue illustration for CosoryxC

Cosoryx

Cosoryx spp.

An extinct pronghorn relative recognised by branched horn cores, with uncertain sheath shape and species boundaries.

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Catalogue illustration for CricetodonC

Cricetodon

Cricetodon spp.

A Miocene cricetodontine rodent whose tooth series reveal multiple lineages.

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Catalogue illustration for CynodontiaC

Cynodontia

Cynodontia Owen, 1861

A mammal-line clade of therapsid synapsids, distinct from dinosaurs and represented today by mammals.

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Catalogue illustration for CyriacotheriumC

Cyriacotherium

Cyriacotherium argyreum; C. psamminum

A small Paleocene North American mammal whose unusual teeth prompted competing ideas about its relationships.

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D

Catalogue illustration for DeinotheriumD

Deinotherium

Deinotherium

A giant proboscidean with tusks curving down from the lower jaw, known from Africa and Eurasia.

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Catalogue illustration for DesmatolagusD

Desmatolagus

Desmatolagus spp.

An Asian early lagomorph whose wear-sensitive teeth and disputed family placement complicate its history.

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Catalogue illustration for DesmostylusD

Desmostylus

Desmostylus hesperus

A Miocene coastal mammal with column-built molars and articulated skeletons from Japan.

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Catalogue illustration for DiacodexisD

Diacodexis

Diacodexis spp.

An early Eocene artiodactyl with a nearly complete skeleton and a specialized ankle.

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Catalogue illustration for DiceratheriumD

Diceratherium

Diceratherium armatum

A North American rhinoceros with paired nasal horn bases and a taxonomic record refined by repeated revisions.

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Catalogue illustration for DidelphodonD

Didelphodon

Didelphodon vorax Marsh, 1889

A Late Cretaceous metatherian known from a near-complete Hell Creek skull, robust jaws and evidence for a varied diet.

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Didymictis

Didymictis spp.

A viverravid carnivoramorphan whose juvenile teeth corrected an early fossil record.

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Catalogue illustration for DidymoconusD

Didymoconus

Didymoconus colgatei; D. berkeyi

An enigmatic Oligocene Asian mammal known from skulls, teeth and a Chinese specimen with robust forelimbs.

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Catalogue illustration for DimetrodonD

Dimetrodon

Dimetrodon

A diverse Early Permian predatory synapsid known for its tall neural spines and differentiated teeth.

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Dinictis

Dinictis felina

A cat-like North American nimravid distinguished from true cats by skull and ear anatomy.

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Catalogue illustration for DioplotheriumD

Dioplotherium

Dioplotherium manigaulti; D. allisoni

A tusked dugongid sirenian known from incomplete fossils in the Americas; the type of D. manigaulti is a premaxillary fragment and tusk.

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Catalogue illustration for DissacusD

Dissacus

Dissacus praenuntius

A mesonychid mammal whose Bighorn Basin teeth reveal feeding changes around the PETM.

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Catalogue illustration for DorcatheriumD

Dorcatherium

Dorcatherium spp.

A widespread Miocene tragulid with M-structured molars, no antlers and enlarged canines in males.

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Catalogue illustration for DorudonD

Dorudon

Dorudon atrox

A five-metre Eocene whale from Fayum with flippers, reduced hind limbs and a rich fossil record at Wadi Al Hitan.

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Catalogue illustration for DusisirenD

Dusisiren

Dusisiren spp.

Miocene North Pacific sea cows whose California and Japanese fossils preserve changes in teeth and forelimbs.

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Catalogue illustration for DviniaD

Dvinia

Dvinia prima

A small Late Permian cynodont from northern Russia, known from skulls with a secondary palate and complex postcanine teeth.

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E

Catalogue illustration for EctocionE

Ectocion

Ectocion spp.

A phenacodontid whose fossil teeth trace body-size and shape patterns across early Cenozoic warming.

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Catalogue illustration for EctypodusE

Ectypodus

Ectypodus spp.

A multituberculate known mainly from isolated teeth, with rare skull material and an Arctic Canadian occurrence.

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Catalogue illustration for ElasmotheriumE

Elasmotherium

Elasmotherium sibiricum Fischer, 1808

A giant Pleistocene Eurasian rhinoceros with a frontal skull boss and high-crowned grazing teeth.

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Catalogue illustration for EleutherocercusE

Eleutherocercus

Eleutherocercus setifer; E. solidus; E. antiquus

A South American glyptodont studied from armour, skull and tail fossils.

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Catalogue illustration for EllipsodonE

Ellipsodon

Ellipsodon; historical assignments include E. aquilonius and E. priscus

A Paleocene North American mammal known from jaws and teeth, with species names repeatedly transferred or synonymized.

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Catalogue illustration for EntelodonE

Entelodon

Entelodon

A long-legged Eurasian entelodont with an enormous head and evidence for an omnivorous diet.

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Catalogue illustration for EntelodontsE

Entelodonts

Entelodontidae

An extinct family of large artiodactyls known from distinctive skulls, powerful jaws and evidence for varied diets.

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Catalogue illustration for EohyraxE

Eohyrax

Eohyrax rusticus; E. isotemnoides and relatives

An Eocene South American typothere known mainly from teeth; its species roster and family placement have been revised.

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Catalogue illustration for EomorphippusE

Eomorphippus

Eomorphippus obscurus; E. bondi; E. neilopdykei

A South American notohippid traced through teeth, jaws and revised fossil layers.

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Catalogue illustration for EosclerocalyptusE

Eosclerocalyptus

Eosclerocalyptus lineatus, E. proximus and E. tapinocephalus

Argentine glyptodonts of the Late Miocene and Early Pliocene, known from carapace plates, tail tubes and limb bones.

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Catalogue illustration for EosirenE

Eosiren

Eosiren libyca; E. stromeri; other assignments vary by revision

Egyptian Eocene–Oligocene sea cows known from skulls, teeth and postcranial remains; E. libyca preserves a strongly reduced pelvis.

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Catalogue illustration for EotheroidesE

Eotheroides

Eotheroides aegyptiacum

An Eocene sea cow genus with an endocast for its type species and more complete skeletons for later congeners.

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Catalogue illustration for EpicyonE

Epicyon

Epicyon saevus, E. haydeni and relatives

A large borophagine canid with massive jaws and premolars capable of processing bone.

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Catalogue illustration for EsthonyxE

Esthonyx

Esthonyx spp.

An early Eocene tillodont with rooted enlarged incisors and a long, species-rich record in Wyoming.

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Catalogue illustration for EucricetodonE

Eucricetodon

Eucricetodon spp.

An Oligocene cricetid known from diverse tooth samples across Europe and Asia.

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Catalogue illustration for EudinocerasE

Eudinoceras

Eudinoceras mongoliensis and five other recognised species

An Asian Eocene coryphodontid pantodont whose taxonomy and stratigraphic range have been revised through dental comparisons.

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Catalogue illustration for EumyarionE

Eumyarion

Eumyarion spp.

A Miocene cricetid whose overlapping molar characters complicate species identification.

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Catalogue illustration for EurhinodelphisE

Eurhinodelphis

Eurhinodelphis spp.

A Miocene toothed whale with an exceptionally elongated, toothless upper rostrum.

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G

Catalogue illustration for GalerixG

Galerix

Galerix spp.

A Miocene galericine known mainly from teeth that document species turnover and regional biochronology.

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Catalogue illustration for GelastopsG

Gelastops

Gelastops parcus and G. joni

A very small Paleocene mammal known from jaws and teeth in Montana and Wyoming, with a fragmentary referred tooth from Colorado.

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Catalogue illustration for GomphotheriumG

Gomphotherium

Gomphotherium angustidens and relatives

A widespread Miocene proboscidean with an elongated lower jaw, upper and lower tusks, and diets that varied among populations.

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Graecopithecus

Graecopithecus freybergi

Late Miocene eastern Mediterranean ape known from a damaged jaw and one premolar, with disputed hominin affinities.

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Catalogue illustration for GranastrapotheriumG

Granastrapotherium

Granastrapotherium snorki

A giant middle Miocene astrapothere known chiefly from a Colombian mandible and tusks, with an uncertain partial-skull referral from Peru.

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H

Catalogue illustration for HaplaletesH

Haplaletes

Haplaletes disceptatrix, H. pelicatus; H. serior is disputed

A Paleocene North American mammal known from jaws and isolated teeth, with changing species limits and a debated referral of H. serior.

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Catalogue illustration for HaplomylusH

Haplomylus

Haplomylus spp.

A small Paleogene mammal whose limb fossils informed a debated hypothesis about elephant-shrew origins.

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Catalogue illustration for HegetotheriumH

Hegetotherium

Hegetotherium mirabile; H. cerdasensis; H. novum

A Miocene South American notoungulate with distinctive incisors and a revised, still debated species history.

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Catalogue illustration for HemihegetotheriumH

Hemihegetotherium

Hemihegetotherium achataleptum, H. torresi, H. tantillum and H. trilobus

A Miocene South American hegetotheriid known mainly from teeth and skulls, with rarer limb fossils and a revised species history.

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Henricosbornia

Henricosbornia spp.

An early South American notoungulate known chiefly from Paleogene teeth and jaw fragments.

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Heosminthus

Heosminthus primiveris

A small fossil dipodoid traced through cheek teeth, with species limits and family placement under review.

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Heptodon

Heptodon spp.

An early Eocene tapiroid whose feet and skull preserve an anatomy unlike the modern tapir form.

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Herpetotherium

Herpetotherium cf. fugax

A metatherian with rare Oligocene evidence from the ear region, endocast and hind limb.

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Hesperocyon

Hesperocyon gregarius Cope, 1873

An early North American canid combining efficient ground travel with retained climbing features.

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Heterohyus

Heterohyus armatus, H. nanus and related species

An Eocene apatemyid known from European jaws and nearly complete Messel skeletons with elongated fingers.

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Catalogue illustration for HomalodotheriumH

Homalodotherium

Homalodotherium cunninghami

A clawed Patagonian notoungulate studied from skulls and limb bones.

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Homo antecessor

Homo antecessor

An early European human from Gran Dolina, known from fragmentary fossils, cut marks and ancient enamel proteins.

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Homo erectus

Homo erectus

A long-lived early human lineage known from Java, Africa and much of Eurasia, with a fully terrestrial body and a fossil record that resists a single tidy origin story.

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Homo ergaster

Homo ergaster

Early African Homo is known from skulls, teeth and a remarkably complete juvenile skeleton, but its species boundary with Homo erectus remains debated.

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Homo floresiensis

Homo floresiensis

A small-bodied island human from Flores, with a long fossil record and unresolved origins.

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Catalogue illustration for Homo georgicusH

Homo georgicus

Homo georgicus

Dmanisi preserves several small-brained early humans in one landscape, offering a rare view of variation during the first dispersals beyond Africa.

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Homo habilis

Homo habilis

Early African human known from a disputed mixture of skull, jaw, hand and limb fossils associated with Oldowan technology.

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Homo heidelbergensis

Homo heidelbergensis

A name founded on one powerful jaw became a label for varied Middle Pleistocene humans, but its geographic and taxonomic limits remain unsettled.

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Homo rudolfensis

Homo rudolfensis

An early human known chiefly from a broad-faced cranium and separate jaws from East Africa.

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Homo sapiens

Homo sapiens

Our species emerged through a long African history: early fossils combine features in different ways, and later populations dispersed and mixed with other humans.

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Homogalax

Homogalax protapirinus

An early Eocene perissodactyl whose partial skeleton preserves primitive limb proportions and unequal digit counts.

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Hoplocetus

Hoplocetus ritzi

Miocene physeteroid with an associated German series of teeth and vertebrae; isolated tooth assignments remain debated.

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Huilatherium

Huilatherium pluriplicatum

A Colombian Miocene notoungulate best documented by deciduous teeth; newer Castilletes fossils remain identified only to genus.

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Hyaenodon

Hyaenodon leptorhynchus and relatives

A diverse Eocene to Miocene hyaenodont with rear-positioned shearing molars and species-dependent body size.

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Hyopsodus

Hyopsodus spp.

A widespread Eocene mammal known from changing tooth populations and one skull that raised a debated hearing hypothesis.

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Hypnomys

Hypnomys morpheus

Balearic island dormouse known from cave teeth and articulated skeletons; limb use remains debated.

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Hypohippus

Hypohippus spp.

A Miocene three-toed browsing horse known from early tooth finds and later skeletons.

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Hyrachyus

Hyrachyus modestus

A middle Eocene perissodactyl known from an unusually informative skeleton with four-toed forefeet and three-toed hind feet.

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Hyracodon

Hyracodon spp.

A hornless North American rhinocerotoid whose limb proportions complicate the “running rhino” label.

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Hyracotherium

Hyracotherium leporinum

An English early Eocene perissodactyl whose type skull anchors a genus once applied to many different mammals.

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I

Catalogue illustration for InostranceviaI

Inostrancevia

Inostrancevia alexandri and relatives

A large Late Permian gorgonopsian with blade-like upper canines, known from Russia and later African records.

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Interatherium

Interatherium rodens; I. extensus

A Miocene Patagonian notoungulate whose two accepted species were clarified in a 2023 systematic revision.

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Ischyromys

Ischyromys spp.

An early rodent with partial skeletons, a scanned Oligocene skull and a record of species replacement.

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Isotemnus

Isotemnus primitivus and other referred material

A Patagonian Eocene notoungulate known from teeth, jaws and isolated limb bones, with a cautious astragalus-based mass estimate.

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K

Catalogue illustration for Kabwe 1K

Kabwe 1

African Middle Pleistocene Homo

A robust Zambian cranium dated to about 299,000 years ago, with a debated species assignment.

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Kraglievichia

Kraglievichia paranensis; K. carinatum

A Miocene pampathere identified mainly from the sculpturing of its armor plates.

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L

Catalogue illustration for LabidolemurL

Labidolemur

Labidolemur spp.

A North American apatemyid known from type jaws, distinctive teeth and rare cranial and skeletal material.

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Catalogue illustration for LeptaucheniaL

Leptauchenia

Leptauchenia decora and relatives

An Oligocene oreodont with high-set eyes, inflated ear chambers and a debated open-country ecology.

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Leptocyon

Leptocyon spp.

Small North American canids that document the long stem history of living Caninae.

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Leptomeryx

Leptomeryx spp.

A small early ruminant known from White River teeth, skulls and limb bones.

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Litaletes

Litaletes disjunctus; related species assignments debated

A small Paleocene mammal from Montana whose jaws and molars define its type species and test later referrals.

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Litocherus

Litocherus zygeus, L. notissimus and L. lacunatus

A Paleocene North American mammal known chiefly from teeth, with an unusual history of competing classifications and some associated postcranial material.

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Litomylus

Litomylus dissentaneus and related Paleocene species

A North American Paleocene mammal known chiefly from teeth, with changing species synonymies and family placement.

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Livyatan

Livyatan melvillei

A giant Miocene sperm whale known from a powerful skull and functional teeth in both jaws.

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Lystrosaurus

Lystrosaurus

A widespread dicynodont with a keratinous beak and upper tusks, recorded across the Permian–Triassic boundary.

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M

Catalogue illustration for MaxschlosseriaM

Maxschlosseria

Maxschlosseria praeterita

A poorly known Patagonian fossil mammal named from upper molars; its species composition and relationships have been repeatedly reviewed.

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Megacerops

Megacerops coloradensis

A massive late Eocene brontothere with paired skull bosses and a long taxonomic history that includes the name Brontotherium.

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Megacricetodon

Megacricetodon spp.

Miocene cricetid rodents whose teeth track regional lineages and dispersal across Europe.

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Catalogue illustration for MegistotheriumM

Megistotherium

Megistotherium osteothlastes

A giant Libyan hyaenodont known chiefly from a 65-centimetre skull and cutting teeth.

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Meniscoessus

Meniscoessus spp.

A Late Cretaceous multituberculate known from incisors, serrated premolars and molars.

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Merychippus

Merychippus sensu lato

Miocene horses with high-crowned teeth and a three-toed foot, within a branching equid radiation.

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Merychyus

Merychyus spp.

A North American oreodont known from changing species, distinctive cheek teeth and rare partial skeletons.

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Merycoidodon

Merycoidodon

A North American oreodont with selenodont plant-processing teeth and a taxonomic history reshaped by abundant fossils.

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Mesodma

Mesodma spp.

A small multituberculate known from distinctive premolars in Late Cretaceous and Paleocene deposits.

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Mesonyx

Mesonyx obtusidens

A middle Eocene mesonychid known from skulls, teeth and the composite Twin Buttes skeleton.

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Metaxytherium

Metaxytherium spp.

A widespread fossil dugongid with species-specific tusks and skeletons from coastal seas.

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Meteutatus

Meteutatus lagenaformis and related species

A Patagonian Paleogene armadillo known chiefly from armour plates, with uncertain boundaries among several historical species names.

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Miacis

Miacis parvivorus Cope, 1872

The type species of a once broad genus, now placed among early carnivoraforms by anatomical analyses.

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Microdyromys

Microdyromys koenigswaldi

A widespread fossil dormouse genus known mainly from molars whose species boundaries remain under review.

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Microparamys

Microparamys spp.

A tiny early Eocene rodent known from detailed teeth and a still-fragmentary body record.

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Microsyops

Microsyops spp.

A Paleogene mammal whose large fossil sample records cavities and changing tooth form.

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Miniochoerus

Miniochoerus spp.

A North American oreodont known from skulls and skeletons across several late Eocene and Oligocene intervals.

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Mioclaenus

Mioclaenus turgidus Cope, 1881

A Paleocene North American mammal documented mainly by jaw fragments and teeth; its type is associated with Torrejonian strata in New Mexico.

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Miohippus

Miohippus annectens and related forms

An Oligocene three-toed horse whose Oregon teeth exposed the pitfalls of naming species from wear.

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Moeritherium

Moeritherium lyonsi, M. trigodon, M. chehbeurameuri

An early North African proboscidean whose tooth wear and enamel isotopes support a partly aquatic ecology.

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Mylodon

Mylodon darwinii

A South American ground sloth known from Darwin’s type jaw, Patagonian cave remains, preserved hair and plant-bearing coprolites.

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N

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Nakalipithecus

Nakalipithecus nakayamai

Late Miocene Kenyan hominoid known from a partial lower jaw and eleven isolated teeth.

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Neanderthals

Homo neanderthalensis

A distinct human lineage known across Pleistocene Europe and western Asia from fossils, tools and ancient DNA.

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Neoplagiaulax

Neoplagiaulax spp.

A Paleocene multituberculate identified chiefly from incisors, premolars and jaws.

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Nesodon

Nesodon imbricatus

A large Early Miocene toxodontid known from skulls and substantial skeletal remains; body mass is estimated at about 550 kg.

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Nimravids

Nimravidae

An extinct feliform family with several independent forms of enlarged upper canines, separate from true cats.

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Notharctus

Notharctus spp.

An Eocene adapiform known from skulls, limbs and a foot with a probable grooming claw.

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Notopithecus

Notopithecus adapinus Ameghino, 1897

A small Patagonian Eocene notoungulate known from an associated skeleton, growth-related dental variation and revised species names.

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Notostylops

Notostylops murinus Ameghino, 1897

A Patagonian notoungulate known from teeth and a partial skeleton used to study limb function and body-mass estimates.

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Nyctitherium

Nyctitherium spp.

A small Eocene mammal identified from jaws and distinctive teeth.

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O

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Oldfieldthomasia

Oldfieldthomasia furcata and historically named congeners

An Eocene South American notoungulate whose species list and family placement have been repeatedly reassessed from dental fossils.

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Orohippus

Orohippus pumilus

A small Eocene equid whose teeth and multi-toed feet illuminate a branching early horse family tree.

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Orrorin

Orrorin tugenensis

A late Miocene Kenyan hominin represented by femora, jaw fragments and evidence of both bipedal loading and climbing.

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Oxyaena

Oxyaena spp.

An Eocene oxyaenid predator identified by jaws and a specialized shearing dentition.

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P

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Pachyrukhos

Pachyrukhos moyani; P. politus

A small Miocene notoungulate with independently evolved rabbit-like proportions; the modern revision recognizes two species.

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Pakicetus

Pakicetus inachus and related pakicetids

A land-capable early whale recognised by its cetacean ear and artiodactyl ankle anatomy.

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Palaeolagus

Palaeolagus haydeni

An early North American lagomorph with two pairs of upper incisors and skulls studied using micro-CT.

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Palaeoloxodon

Palaeoloxodon antiquus and relatives

Pleistocene straight-tusked elephants ranging from mainland giants to independently evolved island dwarfs.

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Palaeoryctes

Palaeoryctes puercensis and related species

A small North American fossil mammal known mainly from teeth, jaws and fragmentary skulls; its species limits and higher classification remain debated.

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Palaeosinopa

Palaeosinopa spp.

A pantolestid known from teeth and rare skeletons, including a juvenile from Fossil Butte.

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Palaeotherium

Palaeotherium spp.

A diverse European Eocene perissodactyl genus known from skulls, teeth and three-toed limbs.

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Paleotomus

Paleotomus senior; P. milleri; P. carbonensis; P. radagasti; P. junior

Paleocene eutherians known mostly from isolated teeth in western North America; species limits and family placement have shifted between studies.

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Pantolambda

Pantolambda bathmodon Cope, 1882

A Paleocene pantodont from western North America; dental histology offers rare evidence about early mammalian growth.

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Paramys

Paramys Leidy, 1871

Early Eocene rodent with a near-complete skeleton used to study climbing and terrestrial locomotion.

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Paranthropus

Paranthropus

Three African hominins shared unusually large cheek teeth, yet their diets and evolutionary relationships were more varied than the old 'nutcracker' label suggests.

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Paranyctoides

Paranyctoides sternbergi; P. maleficus; P. quadrans and other referred forms

A rare Late Cretaceous eutherian known from teeth and jaw fragments; species limits and Asian referrals remain disputed.

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Pararyctes

Pararyctes pattersoni; P. rutherfordi

Paleocene North American eutherians known mainly from teeth and a partial maxilla; their family placement remains debated.

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Parastrapotherium

Parastrapotherium holmbergi

A large Patagonian astrapothere traced through teeth and jaw fragments.

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Parutaetus

Parutaetus chicoensis; P. chilensis; P. punaensis; P. oliveirai

An early South American armadillo lineage known chiefly from bony carapace plates.

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Peltephilus

Peltephilus spp.

A South American armored xenarthran with bony projections on its head shield, best documented from Oligocene and Miocene fossil localities.

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Peradectes

Peradectes spp.

A Paleogene metatherian whose taxonomy is built chiefly from tiny molars and jaw fragments.

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Pericotoxodon

Pericotoxodon platignathus

A large middle Miocene Colombian toxodontid known from partial skulls, teeth and a femur, with modelled mass estimates and fossil bite traces.

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Periphragnis

Periphragnis harmeri Roth, 1899

A South American notoungulate represented by separate Eocene and Oligocene records in Patagonia and Chile.

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Periptychus

Periptychus carinidens

A stout Paleocene mammal with folded teeth, a robust skull and plantigrade limbs.

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Phenacodaptes

Phenacodaptes sabulosus

A late Paleocene Wyoming mammal studied mainly from its jaws and teeth.

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Phenacodus

Phenacodus primaevus Cope, 1873 and relatives

A five-toed early placental whose skeletons document incipient terrestrial running specialisation.

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Phiomia

Phiomia serridens; P. major

An Oligocene early proboscidean with projecting lower incisors.

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Platybelodon

Platybelodon danovi

A Miocene proboscidean with a broadened lower jaw and paired flattened cutting tusks.

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Plesiadapis

Plesiadapis tricuspidens

A Paleogene plesiadapiform with enlarged incisors, claws and a long tail, near but not demonstrably ancestral to primates.

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Pleurostylodon

Pleurostylodon

An Eocene Patagonian notoungulate known from teeth and partial skeletons, including material that corrected a composite reconstruction.

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Pliohippus

Pliohippus spp.

A Miocene horse genus tracing the gradual, branching evolution of a single-toed limb.

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Procerberus

Procerberus

A small cimolestid mammal known from jaw fragments and teeth around the Cretaceous–Paleogene boundary.

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Proeuphractus

Proeuphractus limpidus

A South American Miocene armadillo whose type species is known from separate armour plates and a rediscovered fixed osteoderm.

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Proeutatus

Proeutatus

A Patagonian Miocene armadillo represented by osteoderms, skulls and limb bones; several historical species names are under review.

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Prohegetotherium

Prohegetotherium sculptum; additional referrals disputed

A late Oligocene hegetotheriid from Argentina, with a type species based on two fragmentary upper jaws.

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Promioclaenus

Promioclaenus acolytus and related species

A Paleocene placental mammal known chiefly from teeth and jaw fragments; its family placement and species boundaries have been revised.

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Propalaehoplophorus

Propalaehoplophorus australis

An early Miocene Patagonian glyptodont known from armor and cranial remains.

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Prostegotherium

Prostegotherium notostylopianum; P. astrifer

An early Eocene armadillo-like cingulate diagnosed mostly from patterned armor plates.

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Protoreodon

Protoreodon spp.

An Eocene oreodont revised through comparisons of skulls, teeth and limbs.

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Protoselene

Protoselene opisthacus, P. ashtoni; some referrals remain uncertain

A Paleocene North American mammal known chiefly from teeth and jaw fragments, with a species list shaped by cautious referrals and revisions.

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Protosiren

Protosiren fraasi; P. sattaensis; P. smithae

Eocene sea cows from Egypt and Pakistan whose partial skeletons preserve hind limbs alongside aquatic adaptations.

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Prototherium

Prototherium spp.

Eocene European sea cows known from several skulls and scattered remains; species boundaries remain under review.

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Prototomus

Prototomus spp.

A small early Eocene hyaenodont whose lost type and later Wyoming fossils complicate its taxonomy.

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Protypotherium

Protypotherium spp.

A South American interatheriid whose Miocene teeth and skulls are reshaping species boundaries.

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Prozaedyus

Prozaedyus

A small South American armadillo known from skulls and armor plates, with a changing species taxonomy and younger fragmentary records.

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Pseudhipparion

Pseudhipparion spp.

A diverse genus of three-toed North American horses with species-specific dental specializations.

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Pseudhyrax

Pseudhyrax eutrachytheroides; P. strangulatus

A South American typothere known chiefly from separate teeth; species boundaries and regional referrals require careful comparison.

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Pseudoplohophorus

Pseudoplohophorus orientalis, P. absolutus

A Uruguayan Late Miocene glyptodont known from an associated partial skeleton, armour and a tail tube; some species assignments remain uncertain.

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Ptilodus

Ptilodus montanus Cope, 1881

A Paleocene multituberculate known for specialised chewing teeth and rare partial skeletons that inform debates about movement.

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Puercolestes

Puercolestes simpsoni

An early Paleocene cimolestid known from a damaged New Mexico skull and provisionally referred Montana teeth.

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Pyrotherium

Pyrotherium romeroi, P. macfaddeni

A large Late Oligocene South American mammal known from incomplete skulls, distinctive teeth and scattered limb bones; its relationships and trunk remain uncertain.

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R

Catalogue illustration for RhynchippusR

Rhynchippus

Rhynchippus equinus; R. pumilus; R. brasiliensis

A Patagonian notoungulate whose skull reveals tooth and middle-ear anatomy.

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S

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Sahelanthropus

Sahelanthropus tchadensis

A roughly seven-million-year-old Chadian ape known for the Toumaï cranium and disputed evidence of bipedalism.

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Scirrotherium

Scirrotherium hondaensis; S. antelucanus

A Miocene pampathere from Colombia and Costa Rica whose skull and armor prompted a taxonomic revision.

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Sciuravus

Sciuravus spp.

An Eocene sciuravid rodent whose skull and teeth complicate the squirrel comparison.

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Sifrhippus

Sifrhippus sandrae

An early equid whose teeth document body-size change during rapid PETM warming.

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Sinclairella

Sinclairella dakotensis; S. simplicidens

A rare apatemyid known from a crushed Eocene skull and scattered teeth, including a late Oligocene Florida species.

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Sinolagomys

Sinolagomys spp.

An early Central Asian pika relative known from a long, tooth-rich Oligocene–Miocene record.

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Stegodon

Stegodon

An African and Asian proboscidean genus with ridged molars, diverse diets and multiple island dwarf lineages.

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Stegotherium

Stegotherium tessellatum, S. notohippidensis, S. tauberi

A Miocene Patagonian armadillo with a long snout, reduced teeth and a debated type series.

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Stenotatus

Stenotatus patagonicus

A South American fossil armadillo identified chiefly by three-ridged movable osteoderms; its type material and species boundaries are debated.

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T

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Teleoceras

Teleoceras major Hatcher, 1894 and relatives

A short-limbed rhinoceros known from many articulated Miocene skeletons at Ashfall in Nebraska.

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Tetonius

Tetonius homunculus

A small early Eocene primate from Wyoming whose teeth and new basin records have reshaped its taxonomic story.

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Therocephalians

Therocephalia

A diverse Permian and Triassic therapsid group known for varied skulls, differentiated teeth and a fossil record across Pangaea.

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Thomashuxleya

Thomashuxleya externa and T. rostrata

A middle-Eocene Patagonian notoungulate known from an associated skeleton that links the skull with major parts of the limbs and girdles.

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Thrinaxodon

Thrinaxodon liorhinus

An Early Triassic cynodont known from articulated fossils and burrow associations in southern Gondwana.

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Thryptacodon

Thryptacodon spp.

A fragmentary early placental known for a possible tooth-comb-like arrangement.

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Thylacoleo

Thylacoleo carnifex

A Pleistocene Australian marsupial predator with shearing premolars, powerful forelimbs and an unusually rich skeletal record.

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Thylacosmilus

Thylacosmilus atrox

A South American sparassodont with ever-growing upper canines and a debated role in hunting and carcass processing.

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Thyrohyrax

Thyrohyrax domorictus and related species

An early Oligocene Fayum hyracoid known from a type mandible, later crania and an enigmatic chamber in some jaws.

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Titanoides

Titanoides primaevus

A large Late Paleocene pantodont known mainly from rare teeth and jaw fragments in North America.

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Trachytherus

Trachytherus spegazzinianus, T. alloxus

A South American late Oligocene notoungulate known especially from skulls and teeth; Salla specimens refined its anatomy while leaving genus limits debated.

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Trigonostylops

Trigonostylops wortmani

A Patagonian Eocene astrapothere known from a notable skull, jaw fragments and isolated teeth.

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Trogosus

Trogosus spp.

A large middle Eocene tillodont known from jaws, skulls, limb bones and CT-based cranial studies.

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Typotheriopsis

Typotheriopsis chasicoensis, T. silveyrai

An Argentine mesotheriid with two principal species, a debated premolar diagnosis and newly reported associated skeletal material.

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U

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Ugandax

Ugandax coryndonae

Early African bovin documented by Pliocene skulls, horn cores and teeth; its precise relationship to living buffalo remains debated.

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Uintatherium

Uintatherium anceps

A heavy North American herbivore with three pairs of skull bosses and long upper canines.

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Ustatochoerus

Ustatochoerus spp.

A late oreodont genus with revised species boundaries and increasingly tall cheek teeth.

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Utaetus

Utaetus buccatus

An Eocene South American armadillo known mainly from osteoderms, with a revised type history and a confirmed occurrence in southern Brazil.

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V

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Vassallia

Vassallia minuta; V. maxima (disputed assignment)

An Argentine pampathere known from jaws and armor; the famous V. maxima skull has been reassigned in later revisions.

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Vetelia

Vetelia puncta, V. perforata and V. gandhii

A Miocene armadillo from Argentina and Chile, known from armour plates and rare skulls; a morphological analysis proposed new relationships.

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Villanyia

Villanyia exilis

Small fossil vole defined from Hungarian molars; revised enamel evidence limits older assignments to the genus.

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Viverravus

Viverravus spp.

A small viverravid known mostly from teeth, with one partial skeleton supporting agile climbing in V. acutus.

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Vulpavus

Vulpavus spp.

An Eocene carnivoraform whose limb bones support climbing ability.

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X

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Xenastrapotherium

Xenastrapotherium christi; X. kraglievichi; X. amazonense; X. chaparralensis; X. aequatorialis

Miocene astrapotheres from tropical South America, known mostly from teeth and jaws; genus limits remain under review.

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Continue through deep time

Explore the Cenozoic world

Place every animal within geological time and the methods used to reconstruct extinct bodies.

A catalogue of many branches

“Ancient mammals” is a practical collection rather than a formal lineage. Ambulocetus and Basilosaurus document different stages of early whale evolution. Entelodon belonged to an extinct cetartiodactyl family, while Arsinoitherium and Uintatherium came from separate herbivore radiations.

The collection therefore compares evolutionary experiments without treating similarity as kinship. Heavy bodies, horn-like skull structures and large canines evolved more than once under different diets and ancestry.

From land to water

Early whales show change as a sequence of functional combinations. Ambulocetus retained a pelvis joined to the spine and limbs capable of bearing weight. Large feet and vertical flexion also made it an effective swimmer in coastal water.

Later Basilosaurus was fully marine. Its pelvis no longer supported walking, hind limbs were tiny and propulsion centred on the rear body and tail. The two genera need not form a direct ancestor-and-descendant pair to document how locomotor systems changed within early Cetacea.

Skulls can mislead body reconstruction

Andrewsarchus is known securely from one enormous skull without a lower jaw or associated body. Wolf-like and entelodont-like bodies are comparative hypotheses. Body mass cannot be calculated reliably until the head-to-body proportion is known.

More complete taxa show why caution matters. Entelodont skulls were disproportionately large relative to their torsos, and Uintatherium carried a long massive head on a barrel-shaped body. A spectacular head is real evidence, but it does not supply missing limb bones.

Horn cores, bosses and canines

Arsinoitherium had two pairs of bony horn cores, while Uintatherium had three pairs of bosses and long upper canines. None is simply a rhinoceros. The similarities arose independently and their outer coverings remain unpreserved.

Display is plausible because these structures became prominent in adults. Combat is possible but harder to demonstrate. Growth series and injuries can test the idea, while colour, horn sheaths and ritual behaviour remain reconstruction.

Teeth reveal feeding mechanics

Teeth do more than label a species carnivore or herbivore. Bilophodont teeth in Arsinoitherium processed vegetation. Low-crowned ridged teeth in Uintatherium fit leaves and shoots. Microscopic wear in Entelodon supports a varied omnivorous diet rather than constant bone crushing.

Direct stomach-region remains provide a still stronger line of evidence for Basilosaurus, including young whales and fish. Every profile distinguishes a possible food from prey or plant matter found with an individual.

Size is measured through assumptions

Skeletal length can remain uncertain when vertebrae are missing. Mass requires another step because muscle, fat and trunk depth do not fossilise. Supporting-bone circumference and volumetric models are useful, but a result should name the specimen and preserve a range.

This is why a one-to-two-tonne estimate for Uintatherium, a broad estimate for Arsinoitherium and the unknown mass of Andrewsarchus are not contradictions. They reflect different completeness and different available measurements.

Reading each reconstruction

Bones, teeth and preserved injuries belong to observation. Movement and diet may be strong inferences when several anatomical and geochemical lines agree. Hair, colour, lips, horn covering, herd size and the exact scene of a hunt usually remain artistic.

Frequently asked questions

What counts as an ancient mammal?

The catalogue covers extinct mammals and close mammaliaform relatives from different branches and intervals. It is a practical collection, not one natural taxonomic group.

Were all ancient mammals small?

No. Early members of some lineages were small, but the Cenozoic produced tonne-scale herbivores and whales more than fifteen metres long.

How do palaeontologists estimate their size?

Complete and partial skeletons provide proportions, while supporting-bone dimensions and volumetric models estimate mass. Results are reported as ranges when material is incomplete.

Why do reconstructions of the same mammal differ?

Bones constrain anatomy, but hair, colour, lips, horn coverings and behaviour are often missing. Different artists may choose different plausible soft tissues.