Caviramus: a Triassic pterosaur known from a jaw

A distinctive jaw, unusual teeth and a disputed taxonomic connection.

Interpretive early pterosaur above a warm Triassic coastal lagoon
Life reconstruction. The body, wings, colour and particular setting are not preserved with the type jaw.

Caviramus schesaplanensis is based on a fragment of a right lower jaw from Late Triassic strata in Switzerland. The preserved teeth have multiple cusps, while parts of the jaw show cavities and a notch. No skull or wing belongs to the type specimen, so a complete head shape and wingspan cannot be read from this fossil.

Quick facts

Scientific nameCaviramus schesaplanensis, Fröbisch and Fröbisch, 2006
HolotypePIMUZ A/III 1225, partial right lower jaw
LocalitySchesaplana, Switzerland
Rock unitAlplihorn Member, Kössen Formation
AgeLate Norian to early Rhaetian, Late Triassic
Preserved jaw lengthAbout 52 mm, partly preserved as an impression
WingspanNot established from the type specimen
Evidence guide

What can the fossils tell us?

One incomplete jaw

PIMUZ A/III 1225 preserves roughly 52 mm of a right lower jaw. It is not associated with a skull or wing bones.

A jaw from Schesaplana

Nadia and Jörg Fröbisch named Caviramus schesaplanensis in 2006 from specimen PIMUZ A/III 1225. The fossil comes from the Alplihorn Member of the Kössen Formation on the western side of Schesaplana. These rocks span the late Norian to early Rhaetian stages near the end of the Triassic. The generic name refers to a cavity and a jaw ramus, whereas the species name recalls the mountain.

The surviving right jaw segment measures about 52 mm. Bone and impressions together trace its outline. Two multicusped teeth remain in place, and the authors identified at least twelve positions. A higher estimate for the full tooth row assumes that spacing continued into the missing portion; those extra teeth were not found. Austriadactylus illustrates the diversity of Triassic pterosaurs, but its anatomy cannot fill the missing parts of this Swiss jaw.

What the jaw reveals

The teeth carry several points rather than simple single tips. A notch marks the lower edge of the jaw, and cavities and depressions occur in the bone. The describing authors suggested that the front of the jaw may have carried a keratinous covering. No such soft tissue survives in the slab.

Tooth shape invites questions about food handling, but no gut contents identify a particular prey. A species-specific head crest, complete body outline or flight style cannot be established from this fragment alone.

Illustrated partial pterosaur jaw in limestone
Educational illustration of a fragmentary fossil, not a photograph or exact rendering of PIMUZ A/III 1225.

The Raeticodactylus question

The Swiss pterosaur Raeticodactylus filisurensis was named from a different, more complete specimen. Similarities between its jaw and that of Caviramus prompted a proposal to place them in the same genus. Fabio Dalla Vecchia's later review considered this plausible while treating their species separately and stressing the need for more material. The original Raeticodactylus description had retained separate genera.

The other specimen's crest and wing measurements therefore should not be presented as direct observations of the Caviramus holotype. Campylognathoides is a later Jurassic pterosaur with another documented anatomy, not a substitute skeleton for Caviramus.

Artistic small pterosaur at a Triassic lagoon
The living animal is reconstructed from comparisons; the type fossil does not preserve its wings, body or colour.

The limits of a life reconstruction

Coastal terrain is consistent with the sedimentary setting, yet the animal's colours, exact wing proportions and behaviour are unknown. Fossil images in this article are educational artwork; the specimen itself is a small, incomplete jaw, not a complete skull or skeleton.

Frequently asked questions

Where was Caviramus discovered?

Its type jaw comes from the Alplihorn Member of the Kössen Formation on Schesaplana in Switzerland.

How many teeth were actually found?

Two multicusped teeth remain in place. At least twelve tooth positions were identified; a longer complete row is an estimate.

Is its wingspan known?

No. The type specimen contains a partial jaw and no wing bones.

Is Raeticodactylus the same animal?

The names were based on different specimens. Similarities have led to a proposed same-genus assignment, but the taxonomic interpretation remains open to revision.