Palaeontological atlas

Ancient amphibian catalogue A to Z

Fossil frogs, gilled lake dwellers and large aquatic predators

The collection includes Devonian and Carboniferous early tetrapods, Mesozoic amphibians and Cenozoic forms.

Cards distinguish modern frogs from temnospondyls and separate preserved skulls and gill structures from restored soft anatomy.

In the catalogue124amphibian profiles
Collection focusWater and landearly tetrapods and later amphibians
Time rangeDevonian to Cenozoicearly tetrapods through later amphibians
Showing 124 of 124 profiles

A

Catalogue illustration for AchelomaA

Acheloma

Acheloma cryptatheria

Early Permian trematopid known from skulls in Texas and Oklahoma, including a CT-studied species from Richards Spur.

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Catalogue illustration for AlbanerpetonA

Albanerpeton

Albanerpeton inexpectatum

Small albanerpetontid known chiefly from diagnostic jaws and skull-roof bones across Cretaceous and Cenozoic sites.

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Catalogue illustration for AmphibamusA

Amphibamus

Amphibamus grandiceps

Small Pennsylvanian temnospondyl from Illinois, distinguished from the recently named Jeanerpeton at Mazon Creek.

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Catalogue illustration for AnaschismaA

Anaschisma

Anaschisma browni

Late Triassic metoposaurid revised from Wyoming type fossils and material assigned from Pennsylvania.

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Catalogue illustration for AngusaurusA

Angusaurus

Angusaurus tsylmensis

Early Triassic Russian trematosaur known chiefly from skull fossils, including a CT-studied referred specimen.

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Catalogue illustration for AnoualerpetonA

Anoualerpeton

Anoualerpeton priscus; A. unicus

A small albanerpetontid known from diagnostic jaw and skull bones in England and Morocco.

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Catalogue illustration for ApachesaurusA

Apachesaurus

Apachesaurus gregorii

Late Triassic metoposaurid from the American Southwest, with juvenile fossils and a putative adult under study.

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Catalogue illustration for AphanerammaA

Aphaneramma

Aphaneramma

A long-snouted Early Triassic trematosaurid adapted to coastal and brackish waters after the end-Permian extinction.

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Catalogue illustration for AralobatrachusA

Aralobatrachus

Aralobatrachus robustus

Cretaceous Uzbek frog known from a type jaw and additional isolated bones of uncertain association.

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Catalogue illustration for ArchegosaurusA

Archegosaurus

Archegosaurus decheni

Lower Permian aquatic temnospondyl from Germany with a well-sampled skull growth series.

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Catalogue illustration for AspidosaurusA

Aspidosaurus

Aspidosaurus chiton

Early Permian Texas dissorophid with a neotype skeleton and a median row of keeled osteoderms.

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B

Catalogue illustration for BatrachichnusB

Batrachichnus

Batrachichnus salamandroides

Paleozoic tetrapod trackway ichnogenus; the exact trackmaker remains uncertain.

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Catalogue illustration for BatrachosauroidesB

Batrachosauroides

Batrachosauroides dissimulans

Cenozoic North American salamander genus known chiefly from a lost type skull and isolated vertebrae.

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Catalogue illustration for BatrachosuchoidesB

Batrachosuchoides

Batrachosuchoides lacer

Fragmentary Early Triassic temnospondyl known from an incomplete Russian skull and comparative Polish material.

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Catalogue illustration for BatrachosuchusB

Batrachosuchus

Batrachosuchus browni; questioned B. watsoni and B. concordi assignments

A Triassic brachyopoid temnospondyl known chiefly from skulls, with debated species boundaries.

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Catalogue illustration for BeelzebufoB

Beelzebufo

Beelzebufo ampinga

A large Late Cretaceous Madagascan frog with a reinforced skull, teeth and bony plates in the skin.

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Catalogue illustration for BeiyanerpetonB

Beiyanerpeton

Beiyanerpeton jianpingensis

Late Jurassic Chinese salamander known from articulated skeletons with bony gill structures.

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Catalogue illustration for BenthosuchusB

Benthosuchus

Benthosuchus sushkini

A large Early Triassic aquatic temnospondyl from Russia, known from numerous long flat skulls and jaws.

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Catalogue illustration for BranchierpetonB

Branchierpeton

Branchierpeton spp.

Small micromelerpetid temnospondyl known from European and Moroccan fossils, including external-gill impressions.

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Catalogue illustration for BranchiosaurusB

Branchiosaurus

Branchiosaurus amblystomus

A small Early Permian temnospondyl whose lake fossils document gills, growth and retention of juvenile features.

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Catalogue illustration for BroiliellusB

Broiliellus

Broiliellus brevis

An Early Permian dissorophid genus known from Texas and New Mexico, including skulls, partial skeletons and dorsal osteoderms.

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Catalogue illustration for BroomistegaB

Broomistega

Broomistega putterilli

Early Triassic South African rhinesuchid preserved beside Thrinaxodon in a burrow, with healed rib fractures.

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C

Catalogue illustration for CacopsC

Cacops

Cacops aspidephorus

Small Early Permian dissorophid with a reinforced dorsal skeleton and three named species from Texas and Oklahoma.

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Catalogue illustration for CeltedensC

Celtedens

Celtedens ibericus

European albanerpetontid known from rare articulated Early Cretaceous skeletons at Las Hoyas.

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Catalogue illustration for ChelotritonC

Chelotriton

Chelotriton paradoxus

Hyperossified European salamandrid known from articulated fossils at Enspel and Randeck maar lakes.

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Catalogue illustration for ChenoprosopusC

Chenoprosopus

Chenoprosopus milleri

An early Permian edopoid known from incomplete skulls and a rare bone histology sample.

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Catalogue illustration for ChomatobatrachusC

Chomatobatrachus

Chomatobatrachus halei

A Tasmanian Early Triassic temnospondyl known from skulls, jaws and growth-related palatal differences.

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Catalogue illustration for ChunerpetonC

Chunerpeton

Chunerpeton tianyiense

Jurassic stem salamander from China, known from articulated skeletons and preserved external gills.

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Catalogue illustration for CochleosaurusC

Cochleosaurus

Cochleosaurus bohemicus; C. florensis

A Carboniferous temnospondyl with Czech growth series and Canadian skulls.

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Catalogue illustration for CyclotosaurusC

Cyclotosaurus

Cyclotosaurus robustus

Late Triassic capitosauroid temnospondyl with a closed otic opening and several distinct skull morphologies.

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Catalogue illustration for CzatkobatrachusC

Czatkobatrachus

Czatkobatrachus polonicus

Early Triassic stem salientian from Poland, known from a small sample of separate pelvic, vertebral and limb bones.

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D

Catalogue illustration for DeltasaurusD

Deltasaurus

Deltasaurus kimberleyensis

Early Triassic Western Australian rhytidosteid known from partial skulls, jaws and two named species.

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Catalogue illustration for DendrerpetonD

Dendrerpeton

Dendrerpeton acadianum

Pennsylvanian temnospondyl from Joggins, known from tree-stump finds and a rare articulated skeleton.

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Catalogue illustration for DiplocaulusD

Diplocaulus

Diplocaulus magnicornis

A Permian lepospondyl known for a broad skull whose lateral projections changed during growth.

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Catalogue illustration for DiploceraspisD

Diploceraspis

Diploceraspis burkei

Permian nectridean known for its boomerang-shaped skull; the function of its lateral extensions remains uncertain.

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Catalogue illustration for DiplovertebronD

Diplovertebron

Diplovertebron punctatum

Fragmentary Late Carboniferous tetrapod from Nýřany, known from lectotype M 522 and incomplete remains.

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Catalogue illustration for DissorophusD

Dissorophus

Dissorophus multicinctus

Early Permian Texas temnospondyl with a distinctive layered arrangement of dorsal armor.

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Catalogue illustration for DoleserpetonD

Doleserpeton

Doleserpeton annectens

Small Early Permian tetrapod from Richards Spur, important to debates about the origins of living amphibians.

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Catalogue illustration for DvinosaurusD

Dvinosaurus

Dvinosaurus campbelli

Russian Permian temnospondyl known from skulls, branchial bones and histological evidence of growth.

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E

Catalogue illustration for EdopsE

Edops

Edops craigi

Early Permian Texas temnospondyl known chiefly from a broad skull and jaws.

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Catalogue illustration for EocaeciliaE

Eocaecilia

Eocaecilia micropodia

Early Jurassic stem caecilian from Arizona with a specialized skull and four greatly reduced limbs.

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Catalogue illustration for EocyclotosaurusE

Eocyclotosaurus

Eocyclotosaurus wellesi

Middle Triassic Arizona temnospondyl known from a well-preserved skull in the Moenkopi Formation.

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Catalogue illustration for EodiscoglossusE

Eodiscoglossus

Eodiscoglossus santonjae

Tiny Early Cretaceous frog from northeastern Spain, known from one articulated but poorly preserved skeleton.

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Catalogue illustration for EopelobatesE

Eopelobates

Eopelobates spp.

Extinct pelobatid frog genus with Cenozoic fossils in North America and Europe.

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Catalogue illustration for EoscapherpetonE

Eoscapherpeton

Eoscapherpeton asiaticum

A Late Cretaceous salamander known from skull CT, growth series and healed injuries.

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Catalogue illustration for EryopsE

Eryops

Eryops megacephalus

Large Early Permian temnospondyl known from skulls, vertebrae, limb bones and a heavily built body plan.

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Catalogue illustration for EryosuchusE

Eryosuchus

Eryosuchus tverdochlebovi

Middle Triassic Russian capitosaur known from fragmentary skulls and jaw material.

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F

Catalogue illustration for FedexiaF

Fedexia

Fedexia striegeli

Late Pennsylvanian Pennsylvania trematopid known from one three-dimensional skull and associated jaws.

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Catalogue illustration for FerganobatrachusF

Ferganobatrachus

Ferganobatrachus riabinini

A Middle Jurassic temnospondyl from Kyrgyzstan, represented by an isolated clavicle, skull bones and vertebral hypocentra.

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G

Catalogue illustration for GerobatrachusG

Gerobatrachus

Gerobatrachus hottoni

Early Permian Texas dissorophoid known from a largely articulated skeleton with a distinctive character mosaic.

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Catalogue illustration for GerrothoraxG

Gerrothorax

Gerrothorax pulcherrimus

A flattened aquatic Triassic temnospondyl with a bony gill apparatus and a suction-feeding skull.

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Catalogue illustration for GlanochthonG

Glanochthon

Glanochthon latirostre; G. angusta; G. lellbachae

Early Permian temnospondyls known from skeletons, a skin impression and evidence of fish prey.

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Catalogue illustration for GobiatesG

Gobiates

Gobiates spp.

Cretaceous Central Asian frog genus known from sculptured skulls and a rare associated postcranial skeleton.

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H

Catalogue illustration for HabrosaurusH

Habrosaurus

Habrosaurus spp.

Cretaceous–Paleocene sirenid genus known chiefly from skulls, teeth and vertebrae in western North America.

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Catalogue illustration for HensonbatrachusH

Hensonbatrachus

Hensonbatrachus kermiti

Campanian frog known from diagnostic isolated skull and pelvic bones in Alberta and Montana.

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I

Catalogue illustration for IchthyostegaI

Ichthyostega

Ichthyostega stensioei

A Late Devonian stem tetrapod with seven hind-foot digits, a finned tail and joints unsuited to an ordinary walking gait.

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Catalogue illustration for IsodectesI

Isodectes

Isodectes obtusus

Early Permian Texas dvinosaurian temnospondyl revised from skull-bearing specimens.

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J

Catalogue illustration for JeholotritonJ

Jeholotriton

Jeholotriton paradoxus

Chinese fossil salamander known from articulated skeletons and a debated Yixian–Daohugou age history.

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K

Catalogue illustration for KaraurusK

Karaurus

Karaurus sharovi

Jurassic Kazakhstan stem salamander known from an unusually complete articulated skeleton.

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Catalogue illustration for KestrosaurusK

Kestrosaurus

Kestrosaurus dreyeri; K. kitchingi

An Early Triassic capitosaurid whose type skull required a careful re-evaluation.

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Catalogue illustration for KokartusK

Kokartus

Kokartus honorarius

Bathonian Kyrgyzstan stem salamander studied through CT skull anatomy and long-bone histology.

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Catalogue illustration for KonzhukoviaK

Konzhukovia

Konzhukovia vetusta; K. tarda; proposed K. sangabrielensis

A Permian temnospondyl known from Ural skulls and a debated Brazilian referral.

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Catalogue illustration for KoolasuchusK

Koolasuchus

Koolasuchus cleelandi

Early Cretaceous chigutisaurid from Victoria, Australia, known from diagnostic jaw fragments and scattered skeletal bones.

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Catalogue illustration for KuruleufeniaK

Kuruleufenia

Kuruleufenia xenopoides

A Patagonian Cretaceous pipid known from a braincase and additional scattered bones.

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L

Catalogue illustration for LatoniaL

Latonia

Latonia nigriventer

Fossil frog genus with a wide Cenozoic range and one living species in the Hula Valley.

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Catalogue illustration for LiaoxitritonL

Liaoxitriton

Liaoxitriton zhongjiani

Early Cretaceous salamander from Liaoning represented by a broad slab-preserved sample.

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Catalogue illustration for LisserpetonL

Lisserpeton

Lisserpeton bairdi

Western North American fossil salamander known chiefly from isolated vertebrae and jaw bones.

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Catalogue illustration for LydekkerinaL

Lydekkerina

Lydekkerina huxleyi

Lower Triassic South African stereospondyl known from diagnostic skulls and rare postcranial remains.

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Catalogue illustration for LyrocephaliscusL

Lyrocephaliscus

Lyrocephaliscus euri

An Early Triassic trematosaur from Spitsbergen, known from skulls and associated skeletal fragments.

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M

Catalogue illustration for MarmorerpetonM

Marmorerpeton

Marmorerpeton wakei

Middle Jurassic stem salamander with fragmentary English species and more complete three-dimensional fossils from Skye.

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Catalogue illustration for MastodonsaurusM

Mastodonsaurus

Mastodonsaurus giganteus

Middle Triassic capitosaur with a broad flat skull, paired lower-jaw tusks and a body length reconstructed from incomplete skeletons.

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Catalogue illustration for MegalotritonM

Megalotriton

Megalotriton filholi; status of M. portisi requires review

A Quercy fossil salamander represented by disarticulated vertebrae and limb bones rather than a complete skeleton.

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Catalogue illustration for MelanerpetonM

Melanerpeton

Melanerpeton tenerum

European branchiosaurid temnospondyl notable for a fossil skin impression preserving a spotted pattern.

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Catalogue illustration for MelosaurusM

Melosaurus

Melosaurus uralensis

Large Permian temnospondyl from Russia with a flattened skull, palatal tusks and a debated species-level family tree.

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Catalogue illustration for MetoposaurusM

Metoposaurus

Metoposaurus diagnosticus

Late Triassic metoposaurid with a broad flat skull, forward-set eyes and preserved lateral-line canals; several European species are recognised.

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Catalogue illustration for MicrobrachisM

Microbrachis

Microbrachis pelikani

Late Carboniferous Nýřany tetrapod studied from a large growth sample with lateral-line and tail-regeneration evidence.

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Catalogue illustration for MicropholisM

Micropholis

Micropholis stowi

Small Early Triassic amphibamiform known from Karoo skeletons, bone histology and Antarctic material.

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Catalogue illustration for MioproteusM

Mioproteus

Mioproteus spp.

European fossil proteid salamanders represented chiefly by isolated vertebrae and jaws.

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N

Catalogue illustration for NigerpetonN

Nigerpeton

Nigerpeton ricqlesi

A late Permian temnospondyl from Niger known from two partial skulls with a distinctive long snout and fang arrangement.

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Catalogue illustration for NotobatrachusN

Notobatrachus

Notobatrachus degiustoi

Middle Jurassic Patagonian frog known from a rich skeletal sample and the oldest documented tadpole.

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O

Catalogue illustration for OnchiodonO

Onchiodon

Onchiodon labyrinthicus

European eryopoid temnospondyl known from growth stages that inform a proposed shift in habitat use.

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Catalogue illustration for OphiderpetonO

Ophiderpeton

Ophiderpeton kirktonense

Rare Scottish Carboniferous aïstopod known from fragmentary remains and a revised East Kirkton age.

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Catalogue illustration for OpisthotritonO

Opisthotriton

Opisthotriton kayi

Western North American batrachosauroidid salamander known chiefly from isolated bones.

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P

Catalogue illustration for PalaeobatrachusP

Palaeobatrachus

Palaeobatrachus spp.

European aquatic frog genus with eighteen recognised species and fossil growth series.

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Catalogue illustration for ParalatoniaP

Paralatonia

Paralatonia transylvanica

Small Maastrichtian frog from the Hațeg Basin known from dissociated microvertebrate remains.

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Catalogue illustration for ParanecturusP

Paranecturus

Paranecturus garbanii

Maastrichtian proteid salamander from Montana known from an atlas and other isolated vertebrae.

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Catalogue illustration for ParioxysP

Parioxys

Parioxys ferricolus

An Early Permian Texas temnospondyl whose redescription clarified skull growth and eucacopine affinities.

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Catalogue illustration for ParotosuchusP

Parotosuchus

Parotosuchus nasutus

Early Triassic capitosaur best documented by skulls from Merkel’s Quarry near Bernburg, Germany.

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Catalogue illustration for PhlegethontiaP

Phlegethontia

Phlegethontia spp.

Limbless Paleozoic aïstopod with a long vertebral column and a fenestrated, internally robust skull.

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Catalogue illustration for PiceoerpetonP

Piceoerpeton

Piceoerpeton spp.

Western North American fossil salamander genus represented by two species and mostly isolated vertebrae.

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Catalogue illustration for PlagioscutumP

Plagioscutum

Plagioscutum spp.

Fragmentary Middle Triassic plagiosaurid genus reported from Russia and Kazakhstan.

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Catalogue illustration for PlagiosternumP

Plagiosternum

Plagiosternum granulosum

Broad-skulled Middle Triassic plagiosaurid temnospondyl recorded at multiple German localities.

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Catalogue illustration for PlagiosuchusP

Plagiosuchus

Plagiosuchus pustuliferus

A Middle Triassic temnospondyl with enlarged skull openings and an inferred suction-feeding system.

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Catalogue illustration for PlatyhystrixP

Platyhystrix

Platyhystrix rugosus

Small dissorophid temnospondyl known for tall back plates made from joined vertebral and dermal bone.

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Catalogue illustration for PlatyoposaurusP

Platyoposaurus

Platyoposaurus stuckenbergi

Middle Permian archegosaurid with a long narrow skull, partial skeletons and bone-growth evidence from eastern Europe.

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Catalogue illustration for PliobatrachusP

Pliobatrachus

Pliobatrachus langhae

A palaeobatrachid frog known from fossils spanning the Late Pliocene to Early Pleistocene in Central and Eastern Europe.

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Catalogue illustration for PrionosuchusP

Prionosuchus

Prionosuchus plummeri

Brazilian Permian archegosaurid with a long narrow snout and a body length extrapolated from incomplete remains.

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Catalogue illustration for ProdesmodonP

Prodesmodon

Prodesmodon copei

Fragmentary North American batrachosauroidid salamander distinguished chiefly by its lower jaw.

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Catalogue illustration for ProsalirusP

Prosalirus

Prosalirus bitis

Early Jurassic frog from Arizona whose pelvis and hind limbs record early saltatory anatomy.

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Catalogue illustration for ProsirenP

Prosiren

Prosiren elinorae

An Early Cretaceous Texas salamander known from dorsal vertebrae, with later misassigned jaws separated as Albanerpeton.

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Q

Catalogue illustration for QinglongtritonQ

Qinglongtriton

Qinglongtriton gangouensis

Late Jurassic Chinese salamander represented by a growth series and CT-studied skulls.

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R

Catalogue illustration for RhinesuchusR

Rhinesuchus

Rhinesuchus whaitsi

Late Permian South African temnospondyl revised as part of the early-diverging Rhinesuchidae.

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S

Catalogue illustration for ScapherpetonS

Scapherpeton

Scapherpeton tectum

Fragmentary North American fossil salamander documented mainly by isolated vertebrae.

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Catalogue illustration for SclerocephalusS

Sclerocephalus

Sclerocephalus haeuseri

Permian German temnospondyl known from a growth series and a rediscovered original type skull.

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Catalogue illustration for SclerothoraxS

Sclerothorax

Sclerothorax hypselonotus

Early Triassic German temnospondyl distinguished by tall trunk neural spines and a broad skull.

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Catalogue illustration for ScotiophryneS

Scotiophryne

Scotiophryne

Late Cretaceous North American fossil frog known from an isolated ilium and referred sculptured skull bones.

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Seminobatrachus

Seminobatrachus boltyschkensis

A small Ukrainian fossil frog represented by fourteen skeletons and debated neotenic traits.

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Siderops

Siderops kehli

Large Early Jurassic Australian temnospondyl represented by an articulated skull and much of the skeleton.

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Stanocephalosaurus

Stanocephalosaurus amenasensis

A Triassic Algerian capitosauroid known from well-preserved skulls, including one examined with micro-CT.

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Syrtosuchus

Syrtosuchus samarensis; S. morkovini

Early Triassic Volga-region benthosuchid known chiefly from incomplete skulls.

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T

Catalogue illustration for ThaumastosaurusT

Thaumastosaurus

Thaumastosaurus bottii and related species

Eocene European frog with a sculptured skull and an inferred close relationship to African pyxicephalids.

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Catalogue illustration for TheatoniusT

Theatonius

Theatonius lancensis

A tiny Late Cretaceous frog known from isolated, toothless skull bones; its family-level placement remains uncertain.

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Catalogue illustration for ThoosuchusT

Thoosuchus

Thoosuchus yakovlevi

Early Triassic Russian temnospondyl known chiefly from compact skulls and aquatic sensory grooves.

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Trematosaurus

Trematosaurus brauni

Early Triassic German trematosaurid known from a large cranial sample and natural casts.

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Catalogue illustration for TriadobatrachusT

Triadobatrachus

Triadobatrachus massinoti

Early Triassic Madagascar stem frog known from one partial skeleton with a long bony tail.

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Triassurus

Triassurus sixtelae

Triassic Kyrgyzstan stem salamander known from two larval skeletons in the Madygen Formation.

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Trimerorhachis

Trimerorhachis insignis and related species

Early Permian aquatic temnospondyl from Texas and New Mexico known from abundant cranial and branchial material.

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Catalogue illustration for TupilakosaurusT

Tupilakosaurus

Tupilakosaurus wetlugensis and T. heilmani

Early Triassic dvinosaur from Russia and Greenland, recognised by its skull and paired vertebral centra.

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V

Catalogue illustration for VieraellaV

Vieraella

Vieraella herbstii

Early Jurassic Patagonian frog known from complementary impressions of one partial skeleton.

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W

Catalogue illustration for WatsonisuchusW

Watsonisuchus

Watsonisuchus magnus

Early Triassic South African temnospondyl anchored by a fragmentary skull and jaw type specimen.

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Catalogue illustration for WestlothianaW

Westlothiana

Westlothiana lizziae

Small East Kirkton tetrapod with a nearly complete skeleton and a disputed position near the origin of amniotes.

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Catalogue illustration for WetlugasaurusW

Wetlugasaurus

Wetlugasaurus

An Early Triassic river predator from eastern Europe with a broad flattened skull and conical teeth.

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X

Catalogue illustration for XenotosuchusX

Xenotosuchus

Xenotosuchus africanus

A South African Triassic temnospondyl known from skulls that reveal growth-related changes.

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Z

Catalogue illustration for ZatrachysZ

Zatrachys

Zatrachys serratus

Early Permian temnospondyl recognised by a broad skull, rear projections and a large opening between the nostrils.

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Continue through deep time

Explore arthropod worlds and fossil evidence

Follow the Palaeozoic environments and preservation processes behind shells, tracks and behavioural assemblages.

One catalogue, several branches

Beelzebufo was a true frog. The other animals are temnospondyls, a diverse extinct radiation that should not be pictured as enlarged modern salamanders.

Water leaves several signals in bone

Lateral-line canals occur in Aphaneramma, Benthosuchus and other aquatic forms. Gerrothorax adds a bony branchial apparatus, while Branchiosaurus is known from growth series and rare soft-tissue evidence among its relatives.

Skulls dominate size and diet estimates

Wetlugasaurus and Benthosuchus are much better known from skulls than complete bodies. Total length, tail outline and exact prey therefore remain less secure than teeth and eye position.

Habitats range from lakes to coasts

The cards include freshwater lakes and rivers, floodplains, seasonal terrestrial margins and coastal marine deposits. “Amphibian” does not imply one identical life cycle across deep time.

Frequently asked questions

Were all ancient amphibians close relatives of modern frogs?

No. Beelzebufo was a frog, while the other animals are temnospondyls from extinct tetrapod branches.

Did every temnospondyl need to return to fresh water to breed?

Reproductive biology varies and is poorly preserved. Eggs and larvae are absent for many genera, including coastal Aphaneramma.

How do palaeontologists recognise aquatic life?

Lateral-line canals, gill-supporting bones, limb proportions, dense skeletons and sedimentary setting provide independent evidence.

Why are body-length estimates often approximate?

Skulls are much more complete than trunks and tails, so total length often depends on proportions taken from related animals.