Klobiodon: a Jurassic pterosaur known from a toothy jaw

A fragmentary jaw with strikingly unequal teeth gives this Jurassic pterosaur its name.

Artistic long-tailed pterosaur over a Middle Jurassic English coast
The living animal is an artistic reconstruction based on its jaw and broader pterosaur comparisons. Its complete head, wings and colour are unknown.

Klobiodon rochei was named in 2018 from a partial lower jaw found in the historic Stonesfield quarries of England. Two large recurved front teeth contrast with two much shorter teeth farther back. Although the fossil had been known for many years under other labels, the new diagnosis applies to this jaw, not to every pterosaur bone once associated with those names. No complete skull or wing is securely linked to the holotype.

Quick facts

Scientific nameKlobiodon rochei, O'Sullivan and Martill, 2018
HolotypeNHMUK PV OR 47991, a partial right lower jaw with four preserved teeth
LocalityStonesfield, Oxfordshire, England
Rock and ageStonesfield Slate Member, Taynton Limestone Formation; Bathonian
Jaw140 mm preserved; around 180 mm total length estimated
TeethTwo large recurved front teeth and two shorter medial teeth
WingspanAround 2 m by comparison, not a measured wing
Evidence guide

What can the fossils tell us?

One diagnostic holotype

NHMUK PV OR 47991 preserves 140 mm of the right lower jaw. No associated upper skull, wing or tail establishes the whole body outline.

An old fossil from Stonesfield

Holotype NHMUK PV OR 47991 is kept at London's Natural History Museum. It came from Stonesfield in Oxfordshire, where layered limestone was quarried for roofing material. The named rock unit is the Stonesfield Slate Member of the Taynton Limestone Formation, dated to the Bathonian stage of the Middle Jurassic.

The preserved right lower jaw is 140 mm long. Comparison suggests an original length of about 180 mm, but its missing rear end was not measured. The species name honours comic artist Nick Roche. Historical identifications included Pterodactylus raptor and association with Rhamphocephalus; these were taxonomic interpretations of incomplete fossils.

A jaw that carries the diagnosis

Its two large anterior teeth curve backward and measure about 26 and 18.7 mm. Two following teeth are far shorter, about 7.9 and 10.8 mm. The contrast, together with the form of the jaw and tooth sockets, distinguished Klobiodon in the 2018 study.

Another jaw fragment, OUM J.28410, has been tentatively referred to the genus, but its missing teeth limit a secure assignment. No skull, wings or tail are attached to the type jaw. Dorygnathus is a useful comparison for Jurassic tooth-bearing pterosaurs, not a template that fills these gaps.

Interpretive rendering of a jaw with two tall anterior teeth and shorter following teeth
The illustration shows the difference in tooth size. It is not a photograph or exact tracing of NHMUK PV OR 47991.

Estimates and feeding proposals

An approximately 2 m wingspan is a comparison with other pterosaurs, not a direct measurement from Klobiodon. Its placement among long-tailed forms is based on available anatomical comparisons, and details of its full body remain unknown.

The long front teeth could have helped seize prey, and fish catching has been proposed. No gut contents or prey remnants accompany this jaw. The image of the living animal therefore adds an inferred body plan, colour and behaviour beyond the fossil evidence.

Historical artistic view of layered limestone quarrying near Stonesfield
The quarry scene is illustrative; the workers, day of discovery and pictured slab are invented.

What the old quarry scene can show

An illustration of limestone working evokes the regional history. It does not document the people, date or exact slab involved in discovery of NHMUK PV OR 47991.

Frequently asked questions

Where was Klobiodon found?

Its type jaw came from the Stonesfield Slate Member near Stonesfield, Oxfordshire, England.

Why was it named only in 2018?

The older jaw had been assigned to other pterosaurs before a new comparison found a distinctive combination of jaw and tooth features.

Is the total jaw length known?

Only 140 mm survive; about 180 mm is a comparative estimate of the missing complete length.

Did it definitely catch fish?

No direct food evidence survives. Fish catching is a feeding interpretation based on the teeth.