Melkamter: an early monofenestratan from Jurassic Patagonia

Fragmentary Patagonian bones pushed the known record of monofenestratan pterosaurs into the Early Jurassic.

Speculative early Jurassic pterosaur above a Patagonian waterway
This living animal is reconstructed; the known fossil lacks complete wings, trunk and tail.

Melkamter pateko was described from a broken but informative skull and a few associated bones at Queso Rallado in Patagonia. Its Early Jurassic age and the combined opening ahead of its orbit place it among the earliest known monofenestratan pterosaurs. The specimen is not a complete animal or proof of the exact moment when the group originated.

Quick facts

Scientific nameMelkamter pateko, Fernandes, Pol and Rauhut, 2024
HolotypeMPEF-PV 11530, partial cranium and associated bones
LocalityQueso Rallado, Chubut Province, Argentina
Rock unitLower Cañadón Asfalto Formation
AgeToarcian, latest Early Jurassic
WingspanNot established from the preserved bones
Evidence guide

What can the fossils tell us?

A split skull

The right side of the skull is divided between a main slab and a counterslab; crushing obscures some details.

A find in Chubut

MPEF-PV 11530 came from Queso Rallado near Cerro Cóndor in Chubut Province, Argentina, and is held at the Museo Paleontológico Egidio Feruglio in Trelew. Alexandra Fernandes, Diego Pol and Oliver Rauhut named the genus and species in 2024. Words from Tehuelche supply the name, with the species epithet also recalling the locality and the fossil's fractured state.

The thin silicified mudstone belongs to the lower Cañadón Asfalto Formation, dated to the Toarcian, the latest stage of the Early Jurassic. In the authors' comparison, the specimen predates the previously oldest known monofenestratan by at least eight and probably ten million years. This compares fossil occurrences, rather than dating the lineage's first ancestor precisely.

What the slab preserves

The holotype contains a partial right cranium and counterslab, two associated teeth, four dorsal vertebrae, one metacarpal that could be from finger I, II or III, and unidentifiable fragments. Computed tomography and ultraviolet light helped the authors inspect details of the crushed material.

Although the cranium retains its front and rear extent, the skull roof, occiput and other walls are missing. Its preserved length from snout tip to quadrate is about 13.1 centimetres. This is a cranial measurement, not body length. No complete wing finger or tail permits direct wingspan or tail-length measurement.

Interpretive fieldwork in Jurassic rocks of Patagonia
A generic fieldwork scene, not a documentary image of the Queso Rallado excavation.

The combined opening ahead of the eye

The external naris and antorbital fenestra meet to form a single nasoantorbital opening. A small remnant of the maxilla's ascending process no longer reaches the bones above it. Two phylogenetic analyses using cranial characters recovered Melkamter within early Monofenestrata but outside true pterodactyloids.

Darwinopterus is a later monofenestratan known from more of its skeleton. Rhamphorhynchus illustrates a different long-tailed pterosaur line. A long tail is plausible for Melkamter, but this individual's tail has not been found. Comparison with another Patagonian taxon from the same formation is limited where their preserved bones do not overlap.

Interpretive comparison of two slab surfaces bearing a pterosaur skull
The painted skull is not an exact copy of holotype MPEF-PV 11530.

What remains unknown

The two teeth establish a toothed animal, but not a specific menu. Colour, body covering, flight performance and preferred habitat cannot be measured from this slab. Living restorations use broader knowledge of early pterosaurs and landscape, while the fossil itself establishes a more limited anatomical and temporal record.

Frequently asked questions

Where was Melkamter discovered?

At Queso Rallado near Cerro Cóndor in Argentina's Chubut Province, in the lower Cañadón Asfalto Formation.

How old is the fossil?

The containing strata are Toarcian, at the end of the Early Jurassic. It was the oldest known monofenestratan occurrence in the 2024 analysis.

Which bones survive?

A partial cranium on a slab and counterslab, two associated teeth, four dorsal vertebrae, a metacarpal and small fragments.

Can its wingspan be measured?

No. The holotype does not preserve the complete wing bones needed for a direct span measurement.