Xericeps: a toothless pterosaur jaw from Morocco

A partial lower jaw preserves diagnostic anatomy, while most of the animal is missing.

Cautious artistic reconstruction of a toothless pterosaur beside a Cretaceous North African river
The toothless front jaw is fossil-based; the remaining head, wings, colour and setting are reconstructed.

Xericeps curvirostris was named for the front of a lower jaw found in the Kem Kem beds of Morocco. The fossil is toothless, curves gently upward and carries distinctive grooves and ridges. It does not preserve the rest of the skull, wings or body. The complete animal on the cover is therefore a cautious artistic reconstruction.

Quick facts

Named speciesXericeps curvirostris, Martill and colleagues, 2018
HolotypeFSAC-KK 10700, anterior lower-jaw symphysis
RepositoryHassan II University collection, Casablanca
LocalityAferdou N'Chaft near Hassi el Begaa, southeastern Morocco
Rock unitKem Kem beds; later assigned to the Ifezouane Formation
AgeApproximately Albian to Cenomanian
GroupAzhdarchoid; narrower placement remains uncertain
WingspanUnknown because no wing bone is preserved
Evidence guide

What can the fossils tell us?

Only the lower-jaw front defines the species

FSAC-KK 10700 lacks the rear skull, neck, trunk and wings. Those parts in a full-animal illustration come from comparison with other pterosaurs.

A single fragment from Kem Kem

David Martill and colleagues established Xericeps curvirostris from FSAC-KK 10700, found by local miners near Aferdou N'Chaft in southeastern Morocco. It is the fused front portion of the lower jaws, held in the Hassan II University collection in Casablanca. No associated upper skull, vertebra or wing belongs to the holotype.

The preserved jaw rises gently, and its surface has neither teeth nor tooth sockets. A central dorsal groove bordered by paired ridges toward the rear and a ventral midline groove form part of the diagnostic combination. The species name refers to curvature, but it does not reveal the entire missing beak or upper jaw.

Comparison without a full skull

The authors compared the mandibular symphysis with other toothless azhdarchoid pterosaurs. Alanqa is another Kem Kem taxon, but sharing a region does not make the fossils one species. A subsequent review discussed another fragment, FSAC-KK 5203, in relation to Xericeps. It remains separate material, not a continuation of the holotype.

A more precise position within azhdarchoids would benefit from more associated bones. The rear of the skull and neck cannot be restored confidently from the jaw front alone. There is no wing bone to measure, so a numerical wingspan for this species would be speculative.

Staged artistic examination of a small fossil lower-jaw fragment
A research scene, not a photograph of FSAC-KK 10700; the actual holotype contains only the front lower jaw.

Strata, setting and diet

The original description placed the jaw in the Kem Kem beds near the transition from Early to Late Cretaceous time. A later stratigraphic review refers the relevant beds to the Ifezouane Formation and gives an approximate Albian-Cenomanian range. The revised rock terminology does not add anatomical parts to the fossil.

Deposits in the area suggest a broad ancient river setting, but the actual place where this individual landed or fed is unknown. A toothless jaw section documents local anatomy rather than stomach contents or identified prey. Apatorhamphus offers another example of how fragmentary Kem Kem jaws require care when reconstructing whole pterosaurs.

Artistic toothless pterosaur flying over a Cretaceous river
The river and complete flying animal are imagined. The holotype establishes only part of the lower jaw, not the wings or colour.

What remains artistic

Body size, soft tissues, wing outline and colour cannot be determined for Xericeps from FSAC-KK 10700. The illustrations present one possible azhdarchoid appearance, with a restrained, nearly straight beak rather than exaggerating the gentle curvature visible in the preserved lower jaw.

Frequently asked questions

What is the Xericeps holotype?

FSAC-KK 10700 is the fused front portion of a toothless lower jaw; the rest of the skeleton was not found with it.

Did it have teeth?

No teeth or sockets are present on the preserved jaw fragment. Missing parts cannot be described with the same confidence.

Can its wingspan be measured?

No. The holotype includes no wing bones, so a complete wingspan would rely on uncertain comparisons.

Where was the fossil found?

It comes from the Kem Kem beds near Aferdou N'Chaft in southeastern Morocco, later assigned in a review to the Ifezouane Formation.